Chroniosaurus Temporal range: Upper Permian | |
---|---|
C. dongusensis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Reptiliomorpha (?) |
Order: | † Chroniosuchia |
Family: | † Chroniosuchidae |
Genus: | † Chroniosaurus Tverdochlebova, 1972 |
Species | |
Chroniosaurus is an extinct genus of chroniosuchid stegocephalian, often considered a reptiliomorph [1] but possibly a stem-tetrapod, [2] from upper Permian (upper Tatarian age) deposits of Novgorod, Orenburg and Vologda Regions, Russia. [3] It was first named by Tverdokhlebova in 1972 and the type species is Chroniosaurus dongusensis. [3]
Its lifestyle is uncertain. An early study suggested a fairly aquatic lifestyle, [4] but its femoral microanatomy [5] and dorsal dermal plates [6] suggest a rather terrestrial lifestyle.
Reptiliomorpha is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians (lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes Homo sapiens, but not Ascaphus truei. Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing Homo sapiens, but not Pipa pipa, Caecilia tentaculata, and Siren lacertina.
Pareiasaurs are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with osteoderms which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct in the Permian–Triassic extinction event.
Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and large armour-like bony plates (osteoderms) that generally distinguish them from the modern soft-bodied lissamphibians.
Caseidae are an extinct family of basal synapsids that lived from the Late Carboniferous to Middle Permian between about 300 and 265 million years ago. Fossils of these animals come from the south-central part of the United States, from various parts of Europe, and possibly from South Africa if the genus Eunotosaurus is indeed a caseid as some authors proposed in 2021. Caseids show great taxonomic and morphological diversity. The most basal taxa were small insectivorous and omnivorous forms that lived mainly in the Upper Carboniferous and Lower Permian, such as Eocasea, Callibrachion, and Martensius. This type of caseid persists until the middle Permian with Phreatophasma and may be Eunotosaurus. During the early Permian, the clade is mainly represented by many species that adopted a herbivorous diet. Some have evolved into gigantic forms that can reach 6–7 metres (20–23 ft) in length, such as Cotylorhynchus hancocki and Alierasaurus ronchii, making them the largest Permian synapsids. Caseids are considered important components of early terrestrial ecosystems in vertebrate history because the numerous herbivorous species in this family are among the first terrestrial tetrapods to occupy the role of primary consumer. The caseids experienced a significant evolutionary radiation at the end of the early Permian, becoming, with the captorhinid eureptiles, the dominant herbivores of terrestrial ecosystems in place of the edaphosaurids and diadectids.
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous (Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous (Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape.
Chroniosuchidae is a family of semi-aquatic tetrapods found in sediments from the upper Permian and the upper Triassic periods, most in Russia. They were generally rather large animals, with long jaws similar to those found in modern crocodiles, and probably lived a similar lifestyle as riverside piscivores and ambush predators. Like all chroniosuchians, they bore extensive osteoderm armour on their backs, possibly as protection against terrestrial predators such as the Permian therapsids and the Triassic rauisuchians.
Chroniosuchus is an extinct genus of chroniosuchid from the upper Permian period. The genus was first named by Vjuschkov in 1957.
Uralerpeton is an extinct genus of chroniosuchid reptiliomorph from upper Permian deposits of Vladimir Region, European Russia. It was first named by V. K. Golubev in 1998, from skull fragments and trunk scutes. The type species is Uralerpeton tverdokhlebovae, it was a large predator with a 50 – 55 cm skull and a total length that probably exceeded 3 m (9.3 ft).
Bystrowiana is an extinct genus of bystrowianid chroniosuchian from upper Permian deposits of Vladimir Region, Russia and Jiyuan, China. Chroniosuchians are often thought to be reptiliomorphs, but some recent phylogenetic analyses suggest instead that they are stem-tetrapods. The genus is named in honour of the Russian paleontologist Alexey Bystrow. It was first described by Vyushkov in 1957 and the type species is Bystrowiana permira. Two species—B. permira and B. sinica—are known.
Kotlassia extinct genus of kotlassiine seymouriamorph from the Late Permian of Russia. The type, and currently only, species is K. prima.
Dvinosaurus is an extinct genus of amphibious temnospondyls localized to regions of western and central Russia during the middle and late Permian, approximately 265-254 million years ago. Its discovery was first noted in 1921 by Russian paleontologist Vladimir Prokhorovich Amalitskii in a posthumously published paper that documents the findings of a site in Russia's Arkhangelsk District. Its name is derived from the proximity of this site to the Northern Dvina River.
Chroniosuchia is a group of tetrapods that lived from the Middle Permian to Late Triassic in what is now Eastern Europe, Kyrgyzstan, China and Germany. Chroniosuchians are often thought to be reptiliomorphs, but some recent phylogenetic analyses suggest instead that they are stem-tetrapods. They were all rather short limbed with a strong tail and elongated snout, somewhat resembling modern crocodiles. The group is traditionally considered to be a suborder or order of labyrinthodonts. Chroniosuchians likely had ecological niches as riverside predators, and may have been outcompeted by semiaquatic true reptiles such as phytosaurs in the late Triassic. Most forms bore a heavy armour of scutes along the back, possibly for protection against land born predators like therapsids, or to strengthen the axial skeleton for terrestrial locomotion. Indeed, femoral microanatomy of Chroniosaurus suggests that it was amphibious to terrestrial.
Madygenerpeton is an extinct genus of chroniosuchid reptiliomorph from the Late Triassic Madygen Formation of Kyrgyzstan. It was first named by paleontologists Rainer R. Schoch, Sebastian Voigt and Michael Buchwitz in 2010 from a nearly complete skull and associated osteoderms. The type species is M. pustulatus.
Suchonica is an extinct genus of chroniosuchid reptiliomorph from upper Permian deposits of Sukhona Formation of Vologda Region, Russia. It was first named by V. K. Golubev in 1999, from the anterior armor scute. The type species is Suchonica vladimiri.
Jarilinus is an extinct genus of chroniosuchid reptiliomorph from upper Permian deposits of Novgorod and Orenburg Region, Russia. It was first named by V. K. Golubev in 1998, from skull fragments and trunk scutes. The type species is Jarilinus mirabilis.
Phaanthosaurus is an extinct genus of basal procolophonid parareptile from early Triassic deposits of Nizhnii Novgorod, Russian Federation. It is known from the holotype PIN 1025/1, a mandible. It was collected from Vetluga River, Spasskoe village and referred to the Vokhmian terrestrial horizon of the Vokhma Formation. It was first named by P. K. Chudinov and B. P. Vjushkov in 1956 and the type species is Phaanthosaurus ignatjevi.
Bystrowianidae is a family of chroniosuchian tetrapods from the Permian and Triassic periods.
Laosuchus is an extinct genus of chroniosuchian known from the Permian-Triassic boundary of Asia. Two species have been named.
This list of fossil amphibians described in 2018 is a list of new taxa of fossil amphibians that were described during the year 2018, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2018.
The Timezgadiouine Formation, sometimes spelled as the Timesgadiouine Formation, is a Triassic geological formation in the Argana Basin of Morocco. It is a succession of red bed sediments spanning from the Olenekian to at least the Carnian, encompassing members T3 to T5 of the Argana Group. It is preceded by the Permian Ikakern Formation and succeeded by the Late Triassic Bigoudine Formation.