Bystrowiana Temporal range: Lopingian to Early Triassic | |
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Bystrowiana permira | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Reptiliomorpha (?) |
Order: | † Chroniosuchia |
Family: | † Bystrowianidae |
Genus: | † Bystrowiana Vyushkov, 1957 |
Species | |
Bystrowiana is an extinct genus of bystrowianid chroniosuchian from upper Permian deposits of Vladimir Region, Russia [1] and Jiyuan, China. [2] Chroniosuchians are often thought to be reptiliomorphs, [3] but some recent phylogenetic analyses suggest instead that they are stem-tetrapods. [4] The genus is named in honour of the Russian paleontologist Alexey Bystrow. It was first described by Vyushkov in 1957 and the type species is Bystrowiana permira. [1] Two species—B. permira and B. sinica—are known. [5]
Bystrowiana is known from a 30 cm skull, which suggests it was a large animal, up to 2.5 m (8.2 ft) in total body length. [6]
Bystrowiana in a cladogram after Novikov (2018) showing internal relationships of bystrowianids based on differences in their osteoderms: [7]
Reptiliomorpha is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians (lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes Homo sapiens, but not Ascaphus truei. Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing Homo sapiens, but not Pipa pipa, Caecilia tentaculata, and Siren lacertina.
Pareiasaurs are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with osteoderms which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct in the Permian–Triassic extinction event.
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous (Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous (Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape.
Chroniosuchidae is a family of semi-aquatic tetrapods found in sediments from the upper Permian and the upper Triassic periods, most in Russia. They were generally rather large animals, with long jaws similar to those found in modern crocodiles, and probably lived a similar lifestyle as riverside piscivores and ambush predators. Like all chroniosuchians, they bore extensive osteoderm armour on their backs, possibly as protection against terrestrial predators such as the Permian therapsids and the Triassic rauisuchians.
Chroniosuchus is an extinct genus of chroniosuchid from the upper Permian period. The genus was first named by Vjuschkov in 1957.
Chroniosaurus is an extinct genus of chroniosuchid stegocephalian, often considered a reptiliomorph but possibly a stem-tetrapod, from upper Permian deposits of Novgorod, Orenburg and Vologda Regions, Russia. It was first named by Tverdokhlebova in 1972 and the type species is Chroniosaurus dongusensis.
Benthosuchus is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length.
Chroniosuchia is a group of tetrapods that lived from the Middle Permian to Late Triassic in what is now Eastern Europe, Kyrgyzstan, China and Germany. Chroniosuchians are often thought to be reptiliomorphs, but some recent phylogenetic analyses suggest instead that they are stem-tetrapods. They were all rather short limbed with a strong tail and elongated snout, somewhat resembling modern crocodiles. The group is traditionally considered to be a suborder or order of labyrinthodonts. Chroniosuchians likely had ecological niches as riverside predators, and may have been outcompeted by semiaquatic true reptiles such as phytosaurs in the late Triassic. Most forms bore a heavy armour of scutes along the back, possibly for protection against land born predators like therapsids, or to strengthen the axial skeleton for terrestrial locomotion. Indeed, femoral microanatomy of Chroniosaurus suggests that it was amphibious to terrestrial.
Luzocephalus is an extinct genus of temnospondyl from the Early Triassic of Russia and Greenland. It is usually regarded as a member of the family Lydekkerinidae, although it has also been placed in the family Trematosauridae.
Madygenerpeton is an extinct genus of chroniosuchid reptiliomorph from the Late Triassic Madygen Formation of Kyrgyzstan. It was first named by paleontologists Rainer R. Schoch, Sebastian Voigt and Michael Buchwitz in 2010 from a nearly complete skull and associated osteoderms. The type species is M. pustulatus.
Bystrowiella is an extinct genus of bystrowianid chroniosuchians from upper Middle Triassic deposits of Kupferzell and Vellberg, northern Baden-Württemberg, Germany.
Axitectum is an extinct genus of bystrowianid chroniosuchians from lower Triassic deposits of Nizhni Novgorod and Kirov Regions, Russia. It was a rather large animal judging by the size of its vertebrae. The back was covered in bands of highly ornamented osteoderm plates, similar to those found in modern crocodiles. The bands overlapped with the next band at the posterior end.
Dromotectum is an extinct genus of bystrowianid chroniosuchians from the Late Permian of China and Early Triassic of Russia. Two species have been named: the type species D. spinosum and the species D. largum. D. spinosum, the first species to be named, comes from Lower Triassic deposits in the Samara Region of European Russia and is known from the holotype PIN 2424/23, which consists of armor scutes, and from PIN 2424/65, 4495/14 and 2252/397. It was found in the Staritskaya Formation of the Rybinskin Horizon and named by I.V. Novikov and M.A. Shishkin in 2000. The generic name means “corridor with hipped vault” + “roof” (tecton), and the specific name means “spinous”. A second species, D. largum, was named by Liu Jun, Xu Li, Jia Song-Hai, Pu Han-Yong, and Liu Xiao-Ling in 2014 from the Shangshihezi Formation near Jiyuan in Henan province, China on the basis of specimen IVPP V 4013.1, a large scute.
Synesuchus is an extinct genus of bystrowianid chroniosuchians from middle Triassic deposits of Komi Republic, northern Fore-Ural of Russia. It is known from the holotype PIN 4466/12, which consists of armor scute and from the referred materials PIN 4466/10, 4466/11, 4466/13 and 4466/14. It was found in the Nadkrasnokamenskaya Formation of the Bukobay Horizon. It was first named by I.V. Novikov and M.A. Shishkin in 2000 and the type species is Synesuchus muravjevi. The generic name comes from Syne, from Bolshaya Synya river, and “crocodile”, and the specific name honors the Russian geologist Ivan Stepanovich Murav'ev.
Vonhuenia is an extinct genus of proterosuchid, a basal archosauriform from the Early Triassic of Russia. Fossils have been found in the Vokhminskaya Formation, along the Vetluga River that are Induan in age, making Vonhuenia one of the earliest archosauriforms.
Bystrowianidae is a family of chroniosuchian tetrapods from the Permian and Triassic periods.
Alveusdectes is an extinct genus of diadectid tetrapod from the Late Permian of China. Like other diadectids, it was a large-bodied terrestrial herbivore capable of eating tough fibrous plant material. It was described in 2015 on the basis of a single partial skull and lower jaw found in the Shangshihezi Formation near the city of Jiyuan in Henan. This skull was found in a layer of the Shangshihezi Formation that dates to about 256 million years ago and contains the remains of many other terrestrial tetrapods including pareiasaurs, chroniosuchians, and therapsids. Alveusdectes is the youngest known diadectid by 16 million years and is also the only diadectid known from Asia. It likely represents a late-surviving lineage of diadectids that radiated eastward from western Laurasia into north China. Diadectids are otherwise absent from eastern Laurasia, which may reflect their low diversity at the time. Diadectids first appeared in the Late Carboniferous and were the first animals to have ever occupied the niche of large-bodied herbivores, allowing them to undergo an evolutionary radiation in the Early Permian. By the Late Permian many other groups of tetrapods had entered that niche, and increased competition among herbivores likely resulted in the eventual extinction of diadectids. Alveusdectes may have been able to persist because the fauna of north China seems to have been isolated from other Laurasian faunas during the Late Permian, meaning that fewer herbivores were competing for the same ecological space.
Laosuchus is an extinct genus of chroniosuchian known from the Permian-Triassic boundary of Asia. Two species have been named.
This list of fossil amphibians described in 2018 is a list of new taxa of fossil amphibians that were described during the year 2018, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2018.
Hassiacoscutum is an extinct genus of bystrowianid chroniosuchian found in the Korbach Fissure of the Werra Formation in Hesse, Germany.