Bystrowiana

Last updated

Bystrowiana
Temporal range: Lopingian to Early Triassic
Bystrowiana permira1DB 2.jpg
Bystrowiana permira
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Reptiliomorpha (?)
Order: Chroniosuchia
Family: Bystrowianidae
Genus: Bystrowiana
Vyushkov, 1957
Species
  • B. permira Vyushkov, 1957 (type)
  • B. sinica Young, 1979

Bystrowiana is an extinct genus of bystrowianid chroniosuchian from upper Permian deposits of Vladimir Region, Russia [1] and Jiyuan, China. [2] Chroniosuchians are often thought to be reptiliomorphs, [3] but some recent phylogenetic analyses suggest instead that they are stem-tetrapods. [4] The genus is named in honour of the Russian paleontologist Alexey Bystrow. It was first described by Vyushkov in 1957 and the type species is Bystrowiana permira. [1] Two species—B. permira and B. sinica—are known. [5]

Bystrowiana is known from a 30 cm skull, which suggests it was a large animal, up to 2.5 m (8.2 ft) in total body length. [6]

Phylogeny

Bystrowiana in a cladogram after Novikov (2018) showing internal relationships of bystrowianids based on differences in their osteoderms: [7]

Bystrowianidae

Related Research Articles

<span class="mw-page-title-main">Dissorophidae</span> Extinct family of temnospondyls

Dissorophidae is an extinct family of medium-sized temnospondyls that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America.

<span class="mw-page-title-main">Chroniosuchidae</span> Extinct family of tetrapodomorphs

Chroniosuchidae is a family of semi-aquatic tetrapods found in sediments from the upper Permian and the upper Triassic periods, most in Russia. They were generally rather large animals, with long jaws similar to those found in modern crocodiles, and probably lived a similar life style as riverside piscivores and ambush predators. Like all Chroniosuchians, they bore extensive osteoderm armour on their backs, possibly as protection against terrestrial predators such as the Permian therapsids and the Triassic Rauisuchians.

<i>Chroniosuchus</i> Extinct genus of tetrapods

Chroniosuchus is an extinct genus of chroniosuchid from the upper Permian period. The genus was first named by Vjuschkov in 1957.

<i>Kotlassia</i> Genus of reptile-like amphibians

Kotlassia extinct genus of kotlassiine seymouriamorph from the Late Permian of Russia. The type, and currently only, species is K. prima.

<i>Benthosuchus</i> Extinct genus of amphibians

Benthosuchus is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length.

<span class="mw-page-title-main">Chroniosuchia</span> Extinct order of tetrapods

Chroniosuchia is a group of tetrapods that lived from the Middle Permian to Late Triassic in what is now Eastern Europe, Kyrgyzstan, China and Germany. Chroniosuchians are often thought to be reptiliomorphs, but some recent phylogenetic analyses suggest instead that they are stem-tetrapods. They were all rather short limbed with a strong tail and elongated snout, somewhat resembling modern crocodiles. The group is traditionally considered to be a suborder or order of labyrinthodonts. Chroniosuchians likely had ecological niches as riverside predators, and may have been outcompeted by semiaquatic true reptiles such as phytosaurs in the late Triassic. Most forms bore a heavy armour of scutes along the back, possibly for protection against land born predators like therapsids, or to strengthen the axial skeleton for terrestrial locomotion. Indeed, femoral microanatomy of Chroniosaurus suggests that it was amphibious to terrestrial.

Luzocephalus is an extinct genus of temnospondyl from the Early Triassic of Russia and Greenland. It is usually regarded as a member of the family Lydekkerinidae, although it has also been placed in the family Trematosauridae.

<i>Bystrowiella</i> Extinct genus of tetrapodomorphs

Bystrowiella is an extinct genus of bystrowianid chroniosuchians from upper Middle Triassic deposits of Kupferzell and Vellberg, northern Baden-Württemberg, Germany.

<i>Axitectum</i> Extinct genus of tetrapodomorphs

Axitectum is an extinct genus of bystrowianid chroniosuchians from lower Triassic deposits of Nizhni Novgorod and Kirov Regions, Russia. It was a rather large animal judging by the size of its vertebrae. The back was covered in bands of highly ornamented osteoderm plates, similar to those found in modern crocodiles. The bands overlapped with the next band at the posterior end.

<i>Dromotectum</i> Extinct genus of tetrapodomorphs

Dromotectum is an extinct genus of bystrowianid chroniosuchians from the Late Permian of China and Early Triassic of Russia. Two species have been named: the type species D. spinosum and the species D. largum. D. spinosum, the first species to be named, comes from Lower Triassic deposits in the Samara Region of European Russia and is known from the holotype PIN 2424/23, which consists of armor scutes, and from PIN 2424/65, 4495/14 and 2252/397. It was found in the Staritskaya Formation of the Rybinskin Horizon and named by I.V. Novikov and M.A. Shishkin in 2000. The generic name means “corridor with hipped vault” + “roof” (tecton), and the specific name means “spinous”. A second species, D. largum, was named by Liu Jun, Xu Li, Jia Song-Hai, Pu Han-Yong, and Liu Xiao-Ling in 2014 from the Shangshihezi Formation near Jiyuan in Henan province, China on the basis of specimen IVPP V 4013.1, a large scute.

Synesuchus is an extinct genus of bystrowianid chroniosuchians from middle Triassic deposits of Komi Republic, northern Fore-Ural of Russia. It is known from the holotype PIN 4466/12, which consists of armor scute and from the referred materials PIN 4466/10, 4466/11, 4466/13 and 4466/14. It was found in the Nadkrasnokamenskaya Formation of the Bukobay Horizon. It was first named by I.V. Novikov and M.A. Shishkin in 2000 and the type species is Synesuchus muravjevi. The generic name comes from Syne, from Bolshaya Synya river, and “crocodile”, and the specific name honors the Russian geologist Ivan Stepanovich Murav'ev.

Vonhuenia is an extinct genus of proterosuchid, a basal archosauriform from the Early Triassic of Russia. Fossils have been found in the Vokhminskaya Formation, along the Vetluga River that are Induan in age, making Vonhuenia one of the earliest archosauriforms.

<span class="mw-page-title-main">Bystrowianidae</span> Extinct family of tetrapodomorphs

Bystrowianidae is a family of chroniosuchian tetrapods from the Permian and Triassic periods.

Jiyuanitectum is an extinct genus of chroniosuchian tetrapod from the Late Permian Shangshihezi Formation of China. It is known from a single bony scute from Jiyuan in Henan province, ascribed to the type species Jiyuanitectum flatum in 2014. Plate-like scutes, which formed armor-like coverings on the backs of chroniosuchians, are the most commonly found chroniosuchian remains. They are also the most informative when it comes to distinguishing between species due to small variations in scute anatomy between different taxa. For example, a shallow groove along the midline of the scute is unique to Jiyuanitectum. The flatness of the scute is another unusual characteristic, giving it the species name flatum. Jiyuanitectum shares several features in common with the chroniosuchians Synesuchus and Bystrowiella, including the upper surface of the scute being covered in ridges that are mostly oriented perpendicular to the midline, and the absence of a bony projection on the front of the scute called the anteromedial articular processes, which is seen in other chroniosuchians. These features suggest that Jiyuanitectum belongs to the family Bystrowianidae. The narrowness of the scute suggests that it may be one of the most basal members of the group.

Alveusdectes is an extinct genus of diadectid tetrapod from the Late Permian of China. Like other diadectids, it was a large-bodied terrestrial herbivore capable of eating tough fibrous plant material. It was described in 2015 on the basis of a single partial skull and lower jaw found in the Shangshihezi Formation near the city of Jiyuan in Henan. This skull was found in a layer of the Shangshihezi Formation that dates to about 256 million years ago and contains the remains of many other terrestrial tetrapods including pareiasaurs, chroniosuchians, and therapsids. Alveusdectes is the youngest known diadectid by 16 million years and is also the only diadectid known from Asia. It likely represents a late-surviving lineage of diadectids that radiated eastward from western Laurasia into north China. Diadectids are otherwise absent from eastern Laurasia, which may reflect their low diversity at the time. Diadectids first appeared in the Late Carboniferous and were the first animals to have ever occupied the niche of large-bodied herbivores, allowing them to undergo an evolutionary radiation in the Early Permian. By the Late Permian many other groups of tetrapods had entered that niche, and increased competition among herbivores likely resulted in the eventual extinction of diadectids. Alveusdectes may have been able to persist because the fauna of north China seems to have been isolated from other Laurasian faunas during the Late Permian, meaning that fewer herbivores were competing for the same ecological space.

Parasumina is an extinct genus of anomodont known from the late Capitanian age at the end of the middle Permian period of European Russia. The type and only species is Parasuminia ivakhnenkoi. It was closely related to Suminia, another Russian anomodont, and was named for its resemblance. Little is known about Parasuminia as the only fossils are of fragmentary pieces of the skull and jaw, but the known remains suggest that its head and jaws were deeper and more robust than those of Suminia, and with shorter, stouter teeth. However, despite these differences they appear to have been similar animals with a similarly complex method of processing vegetation.

<i>Laosuchus</i> Extinct genus of tetrapods

Laosuchus is an extinct genus of chroniosuchian known from the Permian-Triassic boundary of Asia. Two species have been named.

<i>Counillonia</i> Extinct genus of dicynodonts

Counillonia is an extinct genus of dicynodont therapsid from the area of Luang Prabang in Laos, Southeast Asia that lived at around the time of the Permian-Triassic boundary and possibly dates to the earliest Early Triassic. Its type and only known species is C. superoculis. Counillonia was related to the Triassic dicynodonts such as Lystrosaurus and the Kannemeyeriiformes that survived the Permian mass extinction, but it was more closely related to the Permian genus Dicynodon than to either of these lineages. Counillonia may then possibly represent another line of dicynodonts that survived the Permian mass extinction into the Triassic period, depending on its age. The discovery of Counillonia in Laos and its unexpected evolutionary relationships hint at the less well understood geographies of dicynodont diversity across the Permo-Triassic boundary outside of well explored regions like the Karoo Basin in South Africa.

<i>Repelinosaurus</i> Extinct genus of dicynodonts

Repelinosaurus is an extinct genus of dicynodont from the Purple Claystone Formation of Luang Prabang in Laos, Southeast Asia that lived at around the time of the Permian-Triassic boundary and possibly dates to the earliest Early Triassic. Its type and only known species is R. robustus. Repelinosaurus was originally described as the earliest known kannemeyeriiform dicynodont, supporting the idea of a more rapid radiation of the Triassic kannemeyeriiform dicynodonts during the Early Triassic following the Permian mass extinction. However, it may alternatively be more closely related to the Permian Dicynodon. The discovery of a potential early kannemeyeriiform in an understudied locality like Laos highlights the importance of such places in dicynodont research, which has been largely focused on historically important localities such as the Karoo Basin of South Africa.

This list of fossil amphibians described in 2018 is a list of new taxa of fossil amphibians that were described during the year 2018, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2018.

References

  1. 1 2 V. K. Golubev (1998). "Narrow-armored Chroniosuchians (Amphibia, Anthracosauromorpha) from the Late Permian of Eastern Europe" (PDF). Paleontological Journal. 32 (3): 278–287.
  2. Young, C.C. (1979). "A new Late Permian fauna from Jiyuan, Honan". Vertebrata PalAsiatica. 17 (2): 99–113.
  3. Michael Buchwitz; Christian Foth; Ilja Kogan; Sebastian Voigt (2012). "On the use of osteoderm features in a phylogenetic approach on the internal relationships of the Chroniosuchia (Tetrapoda: Reptiliomorpha)". Palaeontology. 55 (3): 623–640. Bibcode:2012Palgy..55..623B. doi: 10.1111/j.1475-4983.2012.01137.x .
  4. Laurin, M. (2010). How Vertebrates Left the Water. Berkeley, California, USA.: University of California Press. ISBN   978-0-520-26647-6.
  5. Florian Witzmann; Rainer R. Schoch; Michael W. Maisch (2008). "A relict basal tetrapod from Germany: first evidence of a Triassic chroniosuchian outside Russia" (PDF). Naturwissenschaften. 95 (1): 67–72. Bibcode:2008NW.....95...67W. doi:10.1007/s00114-007-0291-6. PMID   17653527. S2CID   8161364.
  6. Novikov A.V. (2018). Early Triassic amphibians of Eastern Europe: evolution of dominant groups and peculiarities of changing communities (PDF) (in Russian). Moscow: RAS. p. 162. ISBN   978-5-906906-71-7. Archived (PDF) from the original on 2023-12-08. Retrieved 2024-04-06.