Tshekardocoleidae Temporal range: | |
---|---|
Fossil and life restoration of Moravocoleus permianus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Suborder: | † Protocoleoptera |
Family: | † Tshekardocoleidae Rohdendorf, 1944 |
Genera | |
See text | |
Synonyms [1] | |
|
Tshekardocoleidae is an extinct family of stem group beetles, known from the Permian. They represent some of the earliest known beetles. They first appeared during the Cisuralian, before becoming extinct at the beginning of the Guadalupian. [2] A claimed Jurassic record is doubtful. [1] Like other primitive beetles, they are thought to have been xylophagous. [2] The oldest known beetle, Coleopsis, was originally assigned to this family, but is now assigned to its own family Coleopsidae. [1]
The Tshekardocoleidae are thought to have retained several plesiomorphies (ancestral characters) of Coleoptera in a broad sense. For instance, their elytra are flattened, lack epipleura (outer margins), cover the body loosely, and their tips extend beyond the apex of the abdomen. Their abdomens have a nearly cylindrical shape and are thought to have been flexible, apparently being able to strongly contract and expand. In extant beetles, the elytra are tight-fitting, forming a subelytral space which is absent in Tshekardocoleidae. [3]
Kirejtshuk (2020) included the following genera in Tshekardocoleidae [1]
The genus Uralocoleus had been placed in the family by some authors, but the type species Uralocoleus splendensZalessky 1947 was considered incertae sedis by Kirejtshuk (2020) as a possible beetle or cockroach. The second species placed in the genus "Uralocoleus" ultimus was rejected from placement in Uralocoleus and listed as "genus incertus" ultimusPonomarenko, 2000. [1] The genus Dictyocoleus , described from the Jurassic and containing only the type species Dictyocoleus jurassicusHong, 1982, was originally placed in the family but was excluded by Kirejtshuk (2020). [1]
The Permian is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia.
The telephone-pole beetle is a beetle native to the eastern United States and the only living representative of the otherwise extinct family Micromalthidae. Larvae of the beetle live in decaying wood and can be pests to wooden structures, lending them their common name, the 'telephone-pole beetle.'
The Cisuralian is the first series/epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan and dates between 298.9 ± 0.15 – 272.3 ± 0.5 Ma.
In the geologic timescale, the Artinskian is an age or stage of the Permian. It is a subdivision of the Cisuralian Epoch or Series. The Artinskian likely lasted between 290.1 and 283.5 million years ago (Ma) according to the most recent revision of the International Commission on Stratigraphy (ICS) in 2022. It was preceded by the Sakmarian and followed by the Kungurian.
The most recent understanding of the evolution of insects is based on studies of the following branches of science: molecular biology, insect morphology, paleontology, insect taxonomy, evolution, embryology, bioinformatics and scientific computing. It is estimated that the class of insects originated on Earth about 480 million years ago, in the Ordovician, at about the same time terrestrial plants appeared. Insects are thought to have evolved from a group of crustaceans. The first insects were landbound, but about 400 million years ago in the Devonian period one lineage of insects evolved flight, the first animals to do so. The oldest insect fossil has been proposed to be Rhyniognatha hirsti, estimated to be 400 million years old, but the insect identity of the fossil has been contested. Global climate conditions changed several times during the history of Earth, and along with it the diversity of insects. The Pterygotes underwent a major radiation in the Carboniferous while the Endopterygota underwent another major radiation in the Permian.
Ctenospondylus is an extinct genus of sphenacodontid synapsid
Palaeohatteria is an extinct genus of basal sphenacodonts known from the Early Permian period of Saxony, Germany. It contains a single species, Palaeohatteria longicaudata.
Suksunsky District is an administrative district (raion) of Perm Krai, Russia; one of the thirty-three in the krai. Municipally, it is incorporated as Suksunsky Municipal District. It is located in the southeast of the krai. The area of the district is 1,977 square kilometers (763 sq mi). Its administrative center is the urban locality of Suksun. Population: 20,099 (2010 Russian census); 21,925 (2002 Census); 23,654 (1989 Soviet census). The population of Suksun accounts for 39.9% of the district's total population.
Makowskia is an extinct genus of discosauriscid seymouriamorph known from the early Permian of Boskovice Furrow, in the Czech Republic. It was first named by Jozef Klembara in 2005 and the type species is Makowskia laticephala. The generic name honors Alexander Makowsky for describing the first specimens of discosauriscids from the Boskovice Furrow, and the specific name means “broad” + “head”. Makowskia is known only from one specimen, the holotype SNMZ 26506, a skull and anterior portion of postcranial skeleton. A phylogenetic analysis places Makowskia as the sister taxon to Spinarerpeton.
Vladimir Vasilevich Zherikhin, of the Paleontological Institute, Russian Academy of Sciences, Moscow, was one of the world's leading paleoentomologists and coleopterists. He worked on the palaeontology of the Coleoptera (beetles) and of insects in general, and on the taxonomy of the weevils (Curculionoidea).
The Tambach Formation is an Early Permian-age geologic formation in central Germany. It consists of red to brown-colored sedimentary rocks such as conglomerate, sandstone, and mudstone, and is the oldest portion of the Upper Rotliegend within the Thuringian Forest Basin.
Cummingella is a genus of proetid trilobite in the family Phillipsiidae that lived from the earliest Carboniferous until the last species' extinction in the Middle Permian. Fossils have been found in corresponding marine strata of western Europe, the United Kingdom, and the United States.
Brochocoleus is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020.
The Coyote Butte Limestone (OR085) is a geologic formation in Oregon. It preserves fossils dating back to the Sakmarian to Kungurian stages of the Permian period, spanning an estimated 23 million years. The formation occurs in isolated buttes to the north; Triangulation Hill, and south; type locality and name giver Coyote Butte and Tuckers Butte, on either side of the Grindstone and Twelvemile Creeks in Crook County, Oregon.
Tshekardocoleoidea is a superfamily in the extinct suborder Protocoleoptera that contains the following families:
Permocupedidae is a family of Protocoleopteran stem group beetles. They first appeared during the Early Permian, and were one of the dominant groups of beetles during the Middle Permian. They became rare in the Late Permian, with only one species known from the Triassic, Frankencupes ultimus from the Middle Triassic (Anisian) Röt Formation of Germany. They are thought to have been xylophagous, which is presumed to be the ancestral ecology of beetles.
Sweetognathus is an extinct genus of conodonts in the family Sweetognathidae that evolved at the beginning of the Permian period, in near-equatorial, shallow-water seas.
Ademosynidae is an extinct family of beetles, known from the Late Triassic to Early Cretaceous. There are at least six genera and 11 species in Ademosynidae. Members of the family were small oval beetles, with a length generally smaller than 1 cm. Characteristics of the family include a pronotum without anterior angles and a rounded anterior margin, and elytra with 9–12 punctate striae. The systematic position of the family within the order Coleoptera is currently uncertain: they can be considered early examples of the suborder Polyphaga, or unusual examples of Archostemata.
Taldycupedidae is an extinct family of beetles primarily known from the Permian period.