Arcida Temporal range: | |
---|---|
Anadara from the Pliocene of Cyprus. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Bivalvia |
Subclass: | Pteriomorphia |
Order: | Arcida |
Families | |
11, See text. | |
Synonyms | |
Arcoida |
The Arcida is an extant order of bivalve molluscs. [1] This order dates back to the lower Ordovician period. They are distinguished from related groups, such as the mussels, by having a straight hinge to the shells, and the adductor muscles being of equal size. [2] The duplivincular ligament, taxodont dentition, and a shell microstructure consisting of the outer crossed lamellar and inner complex crossed lamellar layers are defining characters of this order. [3]
Seven families are currently recognised within the order, including the well-known ark clams or ark shells in the family Arcidae.
The order Arcida, as the suborder Arcacea, is included in the order Taxodonta by R.C. Moore, 1952, [4] characterised by simple hinge-line dentition consisting of small, numerous, similar hinge teeth, separate mantle lobes, poorly developed siphons, and filibranch gills.
In 2010, Bieler, Carter & Coan [5] proposed a new classification system for the Bivalvia which combines the taxodont Arcida with the dysodont Limida Mytilida, Ostreida and Pteriida as the Pteriomorphia. Subtaxa included in the Arcida are shown below.
In 2016, the superfamilies of Arcida changed from two to three, with an additional superfamily of fossils only. The new taxonomy is as follows: [1] [5]
The Pteriomorphia comprise a subclass of saltwater clams, marine bivalve molluscs. It contains several major orders, including the Arcida, Ostreida, Pectinida, Limida, Mytilida, and Pteriida. It also contains some extinct and probably basal families, such as the Evyanidae, Colpomyidae, Bakevelliidae, Cassianellidae, and Lithiotidae.
Palaeoheterodonta is a subterclass of bivalve molluscs. It contains the extant orders Unionida and Trigoniida. They are distinguished by having the two halves of the shell be of equal size and shape, but by having the hinge teeth be in a single row, rather than separated into two groups, as they are in the clams and cockles.
Trigonioidea is superfamily of medium-sized saltwater clams, marine bivalve molluscs. Within the fossil record the occurrence of this superfamily is widespread, ranging from the Devonian Period to Recent.
Nuculidae is a family of small saltwater clams in the order Nuculida. Species in this family are commonly known as nut clams.
Mactridae, common name the trough shells or duck clams, is a family of saltwater clams, marine bivalve mollusks in the order Venerida.
Lithophaga, the date mussels, are a genus of medium-sized marine bivalve molluscs in the family Mytilidae. Some of the earliest fossil Lithophaga shells have been found in Mesozoic rocks from the Alps and from Vancouver Island.
Mytilida is an order of marine bivalve molluscs, commonly known as true mussels. There is one extant superfamily, the Mytiloidea, with a single extant family, the Mytilidae.
Anomalodesmata is an superorder of saltwater clams, marine bivalve molluscs. This grouping was formerly recognised as a taxonomic subclass. It is called a superorder in the current World Register of Marine Species, despite having no orders, to parallel it with sister taxon Imparidentia, which does have orders.
Praenuculidae is an extinct family of prehistoric bivalves in the superfamily Nuculoidea. Praenuculidae species lived from the early Ordovician, Arenig stage through the Early Devonian Emsian stage. Praenuculidae fossils are found worldwide, present on every continent except Antarctica. Species in this family are thought to have been sessile, attached to the substrate in shallow infaunal marine water environments, where they formed shells of an aragonite composition. The family Praenuculidae was named by A. Lee McAlester in 1969.
Vesicomyidae is a family of saltwater clams, marine bivalve molluscs in the superfamily Glossoidea.
The Pectinoidea are a superfamily of marine bivalve molluscs, including the scallops and spiny oysters.
Periglypta is a genus of bivalves in the subfamily Venerinae of the family Veneridae.
Protobranchia is a subclass of bivalve molluscs. It contains the extant orders Nuculanida, Nuculida, and Solemyida.
Pectinida is a taxonomic order of large and medium-sized saltwater clams, marine bivalve molluscs, commonly known as scallops and their allies. It is believed that they began evolutionarily in the late Middle Ordovician epoch; many species, of course, are still extant.
Poromyoidea is a superfamily of molluscs. It used to contain only the family Poromyidae, but now it also contains Cetoconchidae Ridewood, 1903, as CetoconchaDall, 1886 was removed from Poromyidae and given its own family, according to the World Register of Marine Species.
Trigoniida is an order of medium-sized saltwater clams, marine bivalve molluscs. Within the fossil record the occurrence of this order is widespread, ranging from the Devonian Period to Recent.
In May 2010, a new taxonomy of the Bivalvia was published in the journal Malacologia. The 2010 taxonomy is known as the Taxonomy of the Bivalvia . The 2010 taxonomy was published as Nomenclator of Bivalve Families with a Classification of Bivalve Families. This was a revised system for classifying bivalve mollusks such as clams, oysters, scallops, mussels and so on. In compiling this new taxonomy, the authors used a variety of phylogenetic information including molecular analysis, anatomical analysis, shell morphology and shell microstructure, as well as bio-geographic, paleobiogeographical and stratigraphic information.
Hinge teeth are part of the anatomical structure of the inner surface of a bivalve shell, i.e. the shell of a bivalve mollusk. Bivalves by definition have two valves, which are joined together by a strong and flexible ligament situated on the hinge line at the dorsal edge of the shell. In life, the shell needs to be able to open slightly to allow the foot and siphons to protrude, and then close again, without the valves moving out of alignment with one another. To make this possible, in most cases the two valves are articulated using an arrangement of structures known as hinge teeth. Like the ligament, the hinge teeth are also situated along the hinge line of the shell, in most cases.
Lucinida is a taxonomic order of saltwater clams, marine bivalve molluscs.
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