Bison latifrons Temporal range: | |
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Bison latifrons skeleton | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Family: | Bovidae |
Subfamily: | Bovinae |
Genus: | Bison |
Species: | †B. latifrons |
Binomial name | |
†Bison latifrons | |
Bison latifrons, also known as the giant bison or long-horned bison, is an extinct species of bison that lived in North America during the Late Pleistocene epoch ranging from southern Canada to Mexico. [2] It is noted for its large body size and its distinctive long horns. [3]
Because only skulls and horns of this species have been found well preserved, the size of B. latifrons is currently not clearly known. Based on leg bones, the mass of B. latifrons has been estimated to be 25-50 percent larger than that of modern B. bison, making it undoubtedly one of the largest-ever ruminants. [4] [5]
The known dimensions of the species are on average larger than any extinct and extant bovids, including both the American bison and the European bison, making it the largest known bovid. Overall, it was probably around 4.75 m (15.6 ft) in length and stood about 2.3 m (7.5 ft) or up to 2.5 m (8.2 ft) [6] tall at the withers. [7] With an estimated weight of 1,250 kg (2,760 lb) [8] [9] to 2,000 kg (4,400 lb), [10] B. latifrons was one of the largest ruminants ever, rivaled in mass only by the modern giraffe and the prehistoric long-horned buffalo Pelorovis . Fossil measurements of B. priscus gigas indicate the species was potentially analogous to B. latifrons both in morphology and habitat selection, attaining similar body sizes and horns that were up to 210 centimeters (83 in) apart. [11]
The horns of B. latifrons measured as great as 213 centimeters (84 in) from tip to tip, compared with 66 cm (26 in) to 90 cm (35 in) in modern Bison bison . [12] [13]
Bison latifrons is thought to have evolved in North America from Bison priscus (sometimes called the steppe bison) another prehistoric species of bison that migrated across the Bering Land Bridge around 195–135,000 years ago, before dispersing southwards around 130,000 years ago. The earliest fossil Bison latifrons, as well as one of the oldest well dated bison specimens in central North America, is from the Snowmass site in Colorado, dating to the Last Interglacial (Sangamonian), around 120,000 years ago. [3] It has been suggested that a population of B. latifrons gave rise to Bison antiquus , the ancestor of modern American bison before 60,000 years ago with the two species co-existing alongside each other. [14]
Bison latifrons ranged across much of North America, extending from southern Mexico to southern Canada, as far west as California and as far east as Florida and the American Midwest. [2] Though it was primarily a grazer, B. latifrons exhibited a high degree of dietary flexibility and adaptability across its range. [15] Isotopic analysis from specimens found in Florida suggests a largely grazing based diet, [16] while isotopic analysis of specimens from Clark Quarry, Georgia, suggest that these specimens had a mixed feeding diet (both browsing and grazing), with their diet varying according to the seasons. [17]
While many studies have suggested that the B. latifrons went extinct around or before 20,000 years ago, a 2022 study reported remains from South Texas dating to around 13,000 years Before Present. [18] Pressure from human hunting may have played a role in the extinction of B. latifrons. [19]
A mammoth is any species of the extinct elephantid genus Mammuthus. They lived from the late Miocene epoch into the Holocene until about 4,000 years ago, with mammoth species at various times inhabiting Africa, Asia, Europe, and North America. Mammoths are distinguished from living elephants by their spirally twisted tusks and in at least some later species, the development of numerous adaptions to living in cold environments, including a thick layer of fur.
Bovines comprise a diverse group of 10 genera of medium to large-sized ungulates, including cattle, bison, African buffalo, water buffalos, and the four-horned and spiral-horned antelopes. The members of this group are classified into loose tribes rather than formal subgroups, as the evolutionary relationships within the groups are still uncertain. General characteristics include cloven hooves and usually at least one of the sexes of a species having true horns. The largest extant bovine is the gaur.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders and over 13 tonnes (29,000 lb) in weight, representing among the largest land mammals ever, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some like Palaeoloxodon falconeri less than 1 metre (3.3 ft) in shoulder height as fully grown adults, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
The steppe bison or steppe wisent is an extinct species of bison. It was widely distributed across the mammoth steppe, ranging from Western Europe to eastern Beringia in North America during the Late Pleistocene. It is ancestral to all North American bison, including ultimately modern American bison. Three chronological subspecies, Bison priscus priscus, Bison priscus mediator, and Bison priscus gigas, have been suggested.
The Columbian mammoth is an extinct species of mammoth that inhabited North America from southern Canada to Costa Rica during the Pleistocene epoch. The Columbian mammoth descended from Eurasian steppe mammoths that colonised North America during the Early Pleistocene around 1.5–1.3 million years ago, and later experienced hybridisation with the woolly mammoth lineage. The Columbian mammoth was among the last mammoth species, and the pygmy mammoths evolved from them on the Channel Islands of California. The closest extant relative of the Columbian and other mammoths is the Asian elephant.
Camelops is an extinct genus of camel that lived in North and Central America from the middle Pliocene to the end of the Pleistocene. It is more closely related to living camels than to lamines, making it a true camel of the Camelini tribe. Its name is derived from the Ancient Greek κάμηλος and ὄψ, i.e. "camel-face". Camelops lived across western North America, ranging from the Pacific Coast to the Great Plains, southwards to Honduras and northwards to Alaska. Camelops became extinct as part of the end-Pleistocene extinction event along with most large mammals across the Americas. The extinctions followed the arrival of humans to the Americas, and evidence has been found indicating that humans butchered Camelops, suggesting that hunting may have been a factor in its extinction.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. One of the largest known elephant species, mature fully grown bulls on average had a shoulder height of 4 metres (13 ft) and a weight of 13 tonnes (29,000 lb). Straight-tusked elephants likely lived very similarly to modern elephants, with herds of adult females and juveniles and solitary adult males. The species was primarily associated with temperate and Mediterranean woodland and forest habitats, flourishing during interglacial periods, when its range would extend across Europe as far north as Great Britain and Denmark and eastwards into Russia, while persisting in southern Europe during glacial periods. Skeletons found in association with stone tools and in one case, a wooden spear, suggest they were scavenged and hunted by early humans, including Homo heidelbergensis and their Neanderthal successors.
Bison antiquus is an extinct species of bison that lived in North America during the Late Pleistocene from over 60,000 years ago until around 10,000 years ago. Bison antiquus was one of the most common large herbivores in North America during this time period. It is the direct ancestor of the living American bison.
Hippopotamus gorgops is an extinct species of the genus Hippopotamus known from remains found in Northern Africa, Eastern Africa, and the Levant. One of the largest hippopotamus species, it first appeared during the late Pliocene-Early Pleistocene, and became extinct during the early Middle Pleistocene.
Pelorovis is an extinct genus of African bovines which existed during the Pleistocene epoch. Originally believed to be a giant member of Caprinae, related to modern sheep, it is now known to be a relative of cattle and buffalos. The best known and type species is Pelorovis oldowayensis, from the Early Pleistocene of Olduvai Gorge, Tanzania, though two others, P. turkanensis and P. howelli, are currently recognised. A fourth, P. praeafricanus, may exist, or it may represent the same species as P. oldowayensis. "Pelorovis" antiquus, from the Late Pleistocene-Holocene, and "P." kaisensis, have since been moved into Syncerus, the same genus as living African buffalo.
Palaeoloxodon namadicus is an extinct species of prehistoric elephant known from the Middle Pleistocene to Late Pleistocene of the Indian subcontinent, and possibly also elsewhere in Asia. The species grew larger than any living elephant, and some authors have suggested it to have been the largest known land mammal based on extrapolation from fragmentary remains, though these estimates are speculative.
Hippopotamus antiquus is an extinct species of the genus Hippopotamus that ranged across Europe during the Early and Middle Pleistocene. It was considerably larger than the living hippopotamus.
Bison occidentalis is an extinct species of bison that lived in North America, from about 11,700 to 5,000 years ago, spanning the end of the Pleistocene to the mid-Holocene.
The Late Pleistocene to the beginning of the Holocene saw the extinction of the majority of the world's megafauna, which resulted in a collapse in faunal density and diversity across the globe. The extinctions during the Late Pleistocene are differentiated from previous extinctions by its extreme size bias towards large animals, and widespread absence of ecological succession to replace these extinct megafaunal species, and the regime shift of previously established faunal relationships and habitats as a consequence. The timing and severity of the extinctions varied by region and are thought to have been driven by varying combinations of human and climatic factors. Human impact on megafauna populations is thought to have been driven by hunting ("overkill"), as well as possibly environmental alteration. The relative importance of human vs climatic factors in the extinctions has been the subject of long-running controversy.
Elephas hysudricus is an extinct elephant species known from the Pleistocene of Asia. It is thought to be ancestral to the living Asian elephant, from which it is distinguished by the molar teeth having a lower crown height and a lower lamellae number. Remains of the species are primarily known from the Indian subcontinent, with the most important remains coming from the Siwalik Hills. The oldest remains of the species in the Siwaliks are placed at around 2.6 million year ago at the beginning of the Early Pleistocene, with the youngest dates in the Siwaliks during the Middle Pleistocene around 0.6 million years ago, though it likely persisted on the subcontinent later than this based on remains found elsewhere.
Arctodus is an extinct genus of short-faced bear that inhabited North America during the Pleistocene. There are two recognized species: the lesser short-faced bear and the giant short-faced bear. Of these species, A. simus was larger, is known from more complete remains, and is considered one of the best known members of North America's extinct Ice Age megafauna. A. pristinus was largely restricted to the Early Pleistocene of the eastern United States, whereas A. simus had a broader range, with most finds being from the Late Pleistocene of the United States, Mexico and Canada. A. simus evolved from A. pristinus, but both species likely overlapped in the Middle Pleistocene. Both species are relatively rare in the fossil record.
The Sangamonian Stage is the term used in North America to designate the Last Interglacial and depending on definition, part of the early Last Glacial Period, corresponding to Marine Isotope Stage 5. While often historically considered equivalent in scope to MIS 5, it is now often used in a more narrow sense to refer to the Last Interglacial only. It preceded the Wisconsinan (Wisconsin) Stage and followed the Illinoian Stage in North America.
Hemiauchenia is a genus of laminoid camelids that evolved in North America in the Miocene period about 10 million years ago. This genus diversified and entered South America in the Late Pliocene about three to two million years ago, as part of the Great American Biotic Interchange. The genus became extinct at the end of the Pleistocene. The monophyly of the genus has been considered questionable, with phylogenetic analyses finding the genus to paraphyletic or polyphyletic, with some species suggested to be more closely related to living lamines than to other Hemiauchenia species.
Syncerus antiquus is an extinct species of buffalo from the Late Pleistocene and Holocene of Africa. It was one of the largest species in its family, potentially weighing up to 2,000 kilograms (4,400 lb). Due to this fact, it is sometimes known as the African giant buffalo. The time of its extinction is of debate; Syncerus antiquus either became extinct at the end of the Late Pleistocene about 12,000 years ago or during the Holocene, some 4,000 years ago.
Praeovibos, also known as the giant muskox, is an extinct genus of bovid that contains a single species, Praeovibos priscus. It is closely related to the living muskox, and is placed as a part of the same tribe Ovibovini.