Carcineretidae Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Malacostraca |
Order: | Decapoda |
Suborder: | Pleocyemata |
Infraorder: | Brachyura |
Superfamily: | Portunoidea |
Family: | † Carcineretidae Beurlen, 1930 |
Genera | |
Carcineretidae is a prehistoric family of heterotrematan crustaceans. They are only known from Cretaceous fossils. These crabs are tentatively placed in the superfamily Portunoidea and resemble the swimming crabs (Portunidae) in having some paddle-shaped pereiopods. But it is not certain that this placement is correct, as the Carcineretidae also show some similarities to the Matutidae of superfamily Leucosioidea and the Goneplacidae of superfamily Xanthoidea. [1]
Their carapace is usually wider than long and of square or ovate shape. Its epibranchial region is bulbous, and it bears at least one transverse ridge in the protogastric, hepatic or branchial region. Unlike in the living Portunoidea, the front (the part of the carapace above the head) is narrow; it bears a few blunt spines in some species. The hind part of the carapace is also narrowed, tapering from just behind the eyes. Around the eyes, also atypical for portunoideans, the carapace makes a wide curve and is beset with spines and notches. [1]
The chelipeds are quite robust – more so than the second to fifth pereiopods – and in some the manus is keeled on the outside. Each "paddle" is formed by the flattened propodus and dactyls of the fourth and/or fifth pereiopods. [1]
At least in males, the first and second sternite are fused, with no suture visible. The suture between sternites 2 and 3 is conspicuous and deep, the one between sternites 3 and 4 incomplete. The eighth sternite is not visible from below. The abdominal of males is narrow, with (probably) completely free somites, and the first to fourth abdominal somite entirely fill the space between the coxae of the fifth pereiopods. [1]
Four genera are recognised in the family Carcineretidae: [2]
Dendrobranchiata is a suborder of decapods, commonly known as prawns. There are 540 extant species in seven families, and a fossil record extending back to the Devonian. They differ from related animals, such as Caridea and Stenopodidea, by the branching form of the gills and by the fact that they do not brood their eggs, but release them directly into the water. They may reach a length of over 330 millimetres (13 in) and a mass of 450 grams (1.0 lb), and are widely fished and farmed for human consumption.
Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen), usually hidden entirely under the thorax. They live in all the world's oceans, in freshwater, and on land, are generally covered with a thick exoskeleton, and have a single pair of pincers. They first appeared during the Jurassic Period.
The Decapoda or decapods are an order of crustaceans within the class Malacostraca, and includes crabs, lobsters, crayfish, shrimp, and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 extant species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp and Anomura including hermit crabs, porcelain crabs, squat lobsters making up the bulk of the remainder. The earliest fossils of the group date to the Devonian.
Squat lobsters are dorsoventrally flattened crustaceans with long tails held curled beneath the cephalothorax. They are found in the two superfamilies Galatheoidea and Chirostyloidea, which form part of the decapod infraorder Anomura, alongside groups including the hermit crabs and mole crabs. They are distributed worldwide in the oceans, and occur from near the surface to deep sea hydrothermal vents, with one species occupying caves above sea level. More than 900 species have been described, in around 60 genera. Some species form dense aggregations, either on the sea floor or in the water column, and a small number are commercially fished.
Anomura is a group of decapod crustaceans, including hermit crabs and others. Although the names of many anomurans include the word crab, all true crabs are in the sister group to the Anomura, the Brachyura.
Raninidae is a family of unusual crabs, sometimes known as "frog crabs", on account of their frog-like appearance. They are taken by most scientists to be quite primitive among the true crabs. They closely resemble the (unrelated) mole crabs, due to parallel evolution or convergent evolution. In both groups, the claws are modified into tools for digging, and the body is a rounded shape that is easy to bury in sand. Unlike most other true crabs, the abdomens of raninids are not curled under the cephalothorax.
Cancer is a genus of marine crabs in the family Cancridae. It includes eight extant species and three extinct species, including familiar crabs of the littoral zone, such as the European edible crab, the Jonah crab and the red rock crab. It is thought to have evolved from related genera in the Pacific Ocean in the Miocene.
The Ocypodidae are a family of semiterrestrial crabs that includes the ghost crabs and fiddler crabs. They are found on tropical and temperate shorelines around the world.
Portunoidea is a superfamily of crabs that includes the family Portunidae, the swimming crabs. Which other crab families are also placed here is a matter of some contention, and may be revised following molecular phylogenetic analyses.
Carpilioidea is a superfamily of crabs containing a single extant family, Carpiliidae and three extinct families. The modern range of the family includes the Indo-Pacific, Western Atlantic and Caribbean Sea. The fossil record of the group extends back at least as far as the Paleocene.
Hexapodidae is a family of crabs, the only family in the superfamily Hexapodoidea. It has traditionally been treated as a subfamily of the family Goneplacidae, and was originally described as a subfamily of Pinnotheridae. Its members can be distinguished from all other true crabs by the reduction of the thorax, such that only seven sternites are exposed, and only four pairs of pereiopods are present. Not counting the enlarged pair of claws, this leaves only six walking legs, from which the type genus Hexapus, and therefore the whole family, takes its name. Some anomuran "crabs", such as porcelain crabs and king crabs also have only four visible pairs of legs. With the exception of Stevea williamsi, from Mexico, all the extant members are found either in the Indo-Pacific oceans, or around the coast of Africa.
Heterotremata is a clade of crabs, comprising those crabs in which the genital openings are on the sternum in females, but on the legs in males. It comprises 68 families in 28 superfamilies.
Actumnus is a genus of crabs in the family Pilumnidae. Alongside the 28 extant species, it has a fossil record extending back into the Miocene.
Pseudozioidea is a superfamily of crabs, formerly treated in the Eriphioidea, Carpilioidea, Xanthoidea, Pilumnoidea and Goneplacoidea. A number of fossils from the Eocene onwards are known from the family Pseudoziidae. Eleven genera are recognised in three families:
Crustaceans may pass through a number of larval and immature stages between hatching from their eggs and reaching their adult form. Each of the stages is separated by a moult, in which the hard exoskeleton is shed to allow the animal to grow. The larvae of crustaceans often bear little resemblance to the adult, and there are still cases where it is not known what larvae will grow into what adults. This is especially true of crustaceans which live as benthic adults, more-so than where the larvae are planktonic, and thereby easily caught.
Anatolikos is a genus of two species of crabs in the family Cancridae. They are recorded from Japan and Taiwan. Two fossil species are known, one from Japan and one from Mexico.
Oregonia is a genus of crabs, comprising two extant species and one fossil species: It is classified in the family Oregoniidae under the spider crab superfamily Majoidea.
Paraleptuca chlorophthalmus, is a common fiddler crab found in the mangroves of East Africa, from Somalia to South Africa, as well as Madagascar and Mauritius. Marsh fiddlers dig burrows in the muddy or sandy banks of salt marshes, which they use to protect themselves from predators, high tide and extreme temperatures. They feed by filtering detritus out of mud, and defend their burrows against other fiddler crabs. Paraleptuca chlorophthalmus is characterised by its red pereiopods and blue and black markings on its carapace.
Palaemonella burnsi is a species of shrimp in the family Palaemonidae, from Maui, Hawaii. This species is closest to Palaemonella lata, which it resembles in the broad scaphocerite in which the lamella overreaches the final tooth, and in the unarmed merus of the second pereiopods. It differs from P. lata in the much longer fused part of the two branches of the upper antennular flagellum, in the relatively much longer fingers and shorter palm of the second legs, in the unarmed carpus of the second legs. It is named after John A. Burns, Governor of Hawaii, for declaring the Ahiki Kinau area a nature reserve.
Heteromysis is a genus of marine mysid crustaceans from the family Mysidae, associated with various shallow-water invertebrates. The name describes differentiation of its pereiopods as possible adaptation to commensal life-style. Heteromysis is one of the largest mysid genera, containing more than 100 species. The genus is distributed globally, but predominantly in tropical and subtropical waters.