Chlynovia

Last updated

Chlynovia
Temporal range: Late Permian
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Clade: Therapsida
Suborder: Therocephalia
Family: Ictidosuchidae
Genus: Chlynovia
Tatarinov 2000
Species
  • C. serridentatusTatarinov 2000 (type)

Chlynovia is an extinct genus of therocephalian therapsids from the Late Permian of Russia. The type species is C. serridentatus, named in 2000. [1] Chlynovia was originally classified within Scaloposauria, a group of therocephalians characterized by their small size and lightly built skulls. Scaloposaurians are no longer recognized as a true grouping, but instead represent the juvenile forms of many types of therocephalians. Chlynovia was placed in the family Perplexisauridae along with Perplexisaurus , but both therocephalians are now placed in the family Ictidosuchidae. [2]

Chlynovia was found in the Urpalov Formation in the Kirov region of Russia. Remains of Chlynovia have been found alongside pareiasaurs and therapsids in the Vanyushonkov Member. [3] These animals make up what is known as the Kotelnich assemblage. [2]

Related Research Articles

Gorgonopsia suborder of synapsids (fossil)

Gorgonopsia is an extinct clade of non-mammalian synapsids from the Permian period. Gorgonopsians were quadrupedal predators with prominent canine teeth, and the largest species were the apex predators of their ecosystems. Like other non-mammalian synapsids, gorgonopsians were once described as "mammal-like reptiles", due to their mix of mammalian and reptilian traits. However, this description is no longer considered accurate as they are not reptiles. Rather, as therapsids, they are closely related to mammals. Gorgonopsian fossils have been found in Russia and Africa.

<i>Cistecephalus</i> Assemblage Zone

The Cistecephalus Assemblage Zone is a tetrapod assemblage zone or biozone found in the Adelaide Subgroup of the Beaufort Group, a majorly fossiliferous and geologically important geological group of the Karoo Supergroup in South Africa. This biozone has outcrops located in the Teekloof Formation north-west of Beaufort West in the Western Cape, in the upper Middleton and lower Balfour Formations respectively from Colesberg of the Northern Cape to east of Graaff-Reinet in the Eastern Cape. The Cistecephalus Assemblage Zone is one of eight biozones found in the Beaufort Group, and is considered to be Late Permian in age.

<i>Viatkogorgon</i> genus of mammals (fossil)

Viatkogorgon is an extinct genus of therapsids of Permian Russia. It was a small animal, with well developed gastralia, short legs, and widespread limbs, suggesting that it may have been a good swimmer.

<i>Theriognathus</i> Genus of mammals (fossil)

Theriognathus is an extinct genus of therocephalian therapsid belonging to the family Whaitsiidae, from South Africa and Tanzania. Theriognathus has been dated as existing during the Late Permian. Although Theriognathus means mammal jaw, the lower jaw is actually made up of several bones as seen in modern reptiles, in contrast to mammals. Theriognathus displayed many different reptilian and mammalian characteristics. For example, Theriognathus had canine teeth like mammals, and a secondary palate, multiple bones in the mandible, and a typical reptilian jaw joint, all characteristics of reptiles. It is speculated that Theriognathus was either carnivorous or omnivorous based on its teeth, and was suited to hunting small prey in undergrowth. This synapsid adopted a sleek profile of a mammalian predator, with a narrow snout and around 1 meter long. Theriognathus is represented by 56 specimens in the fossil record.

Tetracynodon is an extinct genus of therocephalian. Fossils of Tetracynodon have been found in the Karoo Basin of South Africa. Two species are known: the type species T. tenuis from the Late Permian and the species T. darti from the Early Triassic. Both species were small-bodied and probably fed on insects and small vertebrates. Although Tetracynodon is more closely related to mammals than it is to reptiles, its braincase is very primitive and shares more in common with modern amphibians and reptiles than it does with mammals.

<i>Scalopodontes</i> genus of mammals (fossil)

Scalopodontes is an extinct genus of therocephalian which existed in Russia during the Late Permian period. The type species is Scalopodontes kotelnichi. Fossils have been found in the Sokolki Assemblage Zone of the Urpalov Formation.

Bauriidae family of mammals (fossil)

Bauriidae is an extinct family of therocephalian therapsids. Bauriids were the latest-surviving group of therocephalians after the Permian–Triassic extinction event, going extinct in the Middle Triassic. They are among the most advanced eutherocephalians and possess several mammal-like features such as a secondary palate and wide postcanine teeth at the back of the jaws. Unlike other therocephalians, bauriids were herbivorous. They were also smaller than earlier members of the group. Two subfamilies are classified within Bauriidae: Nothogomphodontinae and Bauriinae.

Elph is an extinct genus of dicynodont therapsids from Russia. Four specimens have been found from the Sokolki Assemblage in European Russia, representing a fauna that dates back to the Late Permian. Elph was a small herbivore that lived alongside carnivorous akidnognathids and inostranceviids, as well as larger herbivores like Dicynodon and pareiasaurids. The type species E. borealis was named in 1999. Elph has a short snout and tusks and is closely related to Interpresosaurus and Katumbia.

<i>Purlovia</i> genus of mammals (fossil)

Purlovia is an extinct genus of herbivorous therocephalian therapsids from the Late Permian of Russia. Together with the closely related South African genus Nanictidops, it is a member of the family Nanictidopidae. Fossils have been found from the Tonshayevsky District of Nizhny Novgorod Oblast. The type species of Purlovia, P. maxima, was named in 2011.

Muchia is an extinct genus of therocephalian therapsids from the Late Permian of Russia. Fossils have been found in the Kotelnichsky District of Kirov Oblast. The type species M. microdenta was named in 2011. Muchia is known only from a fragment of the lower jaw, making its classification among therocephalians uncertain. It is thought to be within the subfamily Tetracynodontinae, part of the larger group Baurioidea. Other members of the group such as Tetracynodon and Malasaurus are known from more complete material, and have small elongated skulls. Muchia probably had a similar appearance to these therocephalians.

<i>Perplexisaurus</i> genus of mammals (fossil)

Perplexisaurus is a genus of therocephalian therapsid from the Middle to Late Permian Deltavjatia vjatkensis Assemblage Zone, Vanyushonki Member of the Urpalov Formation of Russia. It was described by L. P. Tatarinov in 1997, and the type species is P. foveatus. A new species, P. lepusculus, was described by M.F. Ivachnenko in 2011, from Russia.

Blattoidealestes is an extinct genus of therocephalian therapsid from the Middle Permian of South Africa. The type species Blattoidealestes gracilis was named by South African paleontologist Lieuwe Dirk Boonstra from the Tapinocephalus Assemblage Zone in 1954. Dating back to the Middle Permian, Blattoidealestes is one of the oldest therocephalians. It is similar in appearance to the small therocephalian Perplexisaurus from Russia, and may be closely related.

Homodontosaurus is an extinct genus of therocephalian therapsids from the Late Permian of South Africa. The type species Homodontosaurus kitchingi was named by South African paleontologist Robert Broom in 1949. Broom based his description on a small skull found in the Cistecephalus Assemblage Zone near Graaff-Reinet. The skull is very small, at about 55 millimetres (2.2 in) long and 20 millimetres (0.79 in) wide. Homodontosaurus has large eye sockets and an elongated snout. The lower jaw is long, thin, and curved. Numerous small teeth line the upper jaw and are long, pointed, and round in cross-section.

Silpholestes is an extinct genus of therocephalian therapsids from the Late Permian of South Africa. The type species Silpholestes jackae was named by South African paleontologist Robert Broom in 1948 from the Cistecephalus Assemblage Zone.

Ictidodraco is an extinct genus of therocephalian therapsids from the Late Permian of South Africa. The type species Ictidodraco longiceps was named by South African paleontologists Robert Broom and John T. Robinson in 1948 from the Cistecephalus Assemblage Zone. Ictidodraco was once classified as a scaloposaurian in the family Silpholestidae. Scaloposauria and Silpholestidae are no longer regarded as valid groups, and Ictidodraco is now classified as a basal member of the clade Baurioidea.

Scalopodon is an extinct genus of therocephalian therapsids from the Late Permian of Russia. The type species Scalopodon tenuisfrons was named in 1999 from the Kotelnichsky District of Kirov Oblast. Scalopodon is known from a single fragmentary holotype specimen including the back of the skull, the left side of the lower jaw and isolated postorbital and prefrontal bones. The skull was found in the Deltavjatia Assemblage Zone, which dates back to the early Wuchiapingian about 260 million years ago. Distinguishing features of Scalopodon include narrow frontal bones and a distinctive sagittal crest along the parietal region at the back of the skull. Scalopodon was originally classified in the family Scaloposauridae, and was the first scaloposaurid found in Russia. More recent studies of therocephalians have found scaloposaurids like Scalopodon to be juvenile forms of larger therocephalians and do not consider Scaloposauridae to be a valid group. Scalopodon and most other scaloposaurids are now classified as basal members of Baurioidea.

<i>Karenites</i> genus of mammals (fossil)

Karenites is an extinct genus of therocephalian therapsids from the Late Permian of Russia. The only species is Karenites ornamentatus, named in 1995. Several fossil specimens are known from the town of Kotelnich in Kirov Oblast.

<i>Hazhenia</i> genus of mammals (fossil)

Hazhenia is an extinct genus of therocephalian therapsids from the Early Triassic of China, of which Hazhenia concava is the only species. Hazhenia was named in 1981 from the Heshanggou Formation in the Ordos Desert of Inner Mongolia. It lived during the Olenekian Age of the Early Triassic, about 247 million years ago. Hazhenia belongs to a group of therocephalians called Baurioidea and possesses many mammal-like features such as cusped teeth and a secondary palate, both of which evolved independently in baurioids. Within Baurioidea it is most closely related to the genus Ordosiodon, which is also known from Inner Mongolia but comes from the slightly younger Ermaying Formation. Both genera were once placed in the family Ordosiidae, but as the name is preoccupied by a family of Cambrian trilobites, it is no longer valid.

<i>Gorynychus</i> genus of mammals (fossil)

Gorynychus is a genus of therapsid from the mid-Permian from Kotelnich, Russia. The genus contains two species, G. masyutinae and G. sundyrensis. It was named after the three-headed dragon Zmey Gorynych from Russian mythology.

<i>Nochnitsa</i> genus of mammals (fossil)

Nochnitsa is a genus of gorgonopsian therapsid from the Kotelnich red beds of Permian Russia. It contains one species, Nochnitsa geminidens. It the most basal known gorgonopsian and among the smallest members of the clade known to date.

References

  1. Tatarinov, L.P. (2000). "New material on Scaloposaurians (Reptilia, Theriodontia) from the Upper Permian of the Kotelnich Locality, Kirov Region". Paleontological Journal. 34 (supplement 2): S187–S202.
  2. 1 2 Golubev, V.K. (2000). "The faunal assemblages of Permian terrestrial vertebrates from Eastern Europe" (PDF). Paleontological Journal. 34 (Suppl. 2): S211–S224.
  3. Kordikova, E.G.; Khlyupi, A.J. (2001). "First evidence of a neonate dentition in pareiasaurs from the Upper Permian of Russia" (PDF). Acta Palaeontologica Polonica. 46 (4): 589–594.