Therocephalia is an extinct suborder of eutheriodont therapsids from the Permian and Triassic. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their teeth, suggest that they were carnivores. Like other non-mammalian synapsids, therocephalians were once described as "mammal-like reptiles". Therocephalia is the group most closely related to the cynodonts, which gave rise to the mammals. This relationship takes evidence in a variety of skeletal features. The phylogeny of therocephalians has been disputed, as the monophyly of the group and the relationships of its members are unclear.
Euchambersia is a genus of therocephalian therapsid that lived during the Late Permian, approximately 255 million years ago, in what is now South Africa. The genus contains a single species, Euchambersia mirabilis, named by paleontologist Robert Broom in 1931 from a skull missing the lower jaws; a second skull, belonging to an immature individual, was later described. It is a member of the family Akidnognathidae, which historically has also been referred by as the synonymous Euchambersiidae.
Moschorhinus is an extinct genus of therocephalian in the family Akidnognathidae, with only one species: M. kitchingi. It was a carnivorous, lion-sized synapsid which has been found in the Late Permian to Early Triassic of the South African Karoo Supergroup. It had a broad, blunt snout which bore long, straight canines. It appears to have replaced the gorgonopsids ecologically, and hunted much like a big cat. While most abundant in the Late Permian, it survived a little after the Permian Extinction, though these Triassic individuals had stunted growth.
Pristerognathus is an extinct genus of therocephalian, known from the late Middle Permian (Capitanian) of South Africa. It lends its name to the Pristerognathus Assemblage Zone of the Beaufort Group of South African geological strata. Pristerognathus was a medium-sized therocephalian with a 25 cm (9.8 in) skull and a total length up to 1.5 m.
Theriognathus is an extinct genus of therocephalian therapsid belonging to the family Whaitsiidae, from South Africa and Tanzania. Theriognathus has been dated as existing during the Late Permian. Although Theriognathus means mammal jaw, the lower jaw is actually made up of several bones as seen in modern reptiles, in contrast to mammals. Theriognathus displayed many different reptilian and mammalian characteristics. For example, Theriognathus had canine teeth like mammals, and a secondary palate, multiple bones in the mandible, and a typical reptilian jaw joint, all characteristics of reptiles. It is speculated that Theriognathus was either carnivorous or omnivorous based on its teeth, and was suited to hunting small prey in undergrowth. This synapsid adopted a sleek profile of a mammalian predator, with a narrow snout and around 1 meter long. Theriognathus is represented by 56 specimens in the fossil record.
Regisaurus is an extinct genus of small carnivorous therocephalian. It is known from a single described species, the type species Regisaurus jacobi, from the Early Triassic Lystrosaurus Assemblage Zone of South Africa, although at least one undescribed species is also known.
Cynosaurus is an extinct genus of cynodonts. Remains have been found from the Dicynodon Assemblage Zone in South Africa. Cynosaurus was first described by Richard Owen in 1876 as Cynosuchus suppostus. Cynosaurus has been found in the late Permian period. Cyno- is derived from the Greek word kyon for dog and –sauros in Greek meaning lizard.
Glanosuchus is a genus of scylacosaurid therocephalian from the Late Permian of South Africa. The type species G. macrops was named by Robert Broom in 1904. Glanosuchus had a middle ear structure that was intermediate between that of early therapsids and mammals. Ridges in the nasal cavity of Glanosuchus suggest it had an at least partially endothermic metabolism similar to modern mammals.
Scymnosaurus is now considered a nomen dubium.
Raranimus is an extinct genus of therapsids of the Middle Permian. It was described in 2009 from a partial skull found in 1998 from the Dashankou locality of the Xidagou Formation, outcropping in the Qilian Mountains of Gansu, China. The genus is the most basal known member of the clade Therapsida, to which the later Mammalia belong.
Baurioidea is a superfamily of therocephalian therapsids. It includes advanced therocephalians such as Regisaurus and Bauria. The superfamily was named by South African paleontologist Robert Broom in 1911. Bauriamorpha, named by D. M. S. Watson and Alfred Romer in 1956, is a junior synonym of Bauriodea.
Choerosaurus is an extinct genus of therocephalian therapsids from the Late Permian of South Africa. The type species Choerosaurus dejageri was named by South African paleontologist Sidney H. Haughton from the Tropidostoma Assemblage Zone in 1929.
Blattoidealestes is an extinct genus of therocephalian therapsid from the Middle Permian of South Africa. The type species Blattoidealestes gracilis was named by South African paleontologist Lieuwe Dirk Boonstra from the Tapinocephalus Assemblage Zone in 1954. Dating back to the Middle Permian, Blattoidealestes is one of the oldest therocephalians. It is similar in appearance to the small therocephalian Perplexisaurus from Russia, and may be closely related.
Silpholestes is an extinct genus of therocephalian therapsids from the Late Permian of South Africa. The type species Silpholestes jackae was named by South African paleontologist Robert Broom in 1948 from the Cistecephalus Assemblage Zone.
Ictidodraco is an extinct genus of therocephalian therapsids from the Late Permian of South Africa. The type species Ictidodraco longiceps was named by South African paleontologists Robert Broom and John T. Robinson in 1948 from the Cistecephalus Assemblage Zone. Ictidodraco was once classified as a scaloposaurian in the family Silpholestidae. Scaloposauria and Silpholestidae are no longer regarded as valid groups, and Ictidodraco is now classified as a basal member of the clade Baurioidea.
Scalopodon is an extinct genus of therocephalian therapsids from the Late Permian of Russia. The type species Scalopodon tenuisfrons was named in 1999 from the Kotelnichsky District of Kirov Oblast. Scalopodon is known from a single fragmentary holotype specimen including the back of the skull, the left side of the lower jaw and isolated postorbital and prefrontal bones. The skull was found in the Deltavjatia Assemblage Zone, which dates back to the early Wuchiapingian about 260 million years ago. Distinguishing features of Scalopodon include narrow frontal bones and a distinctive sagittal crest along the parietal region at the back of the skull. Scalopodon was originally classified in the family Scaloposauridae, and was the first scaloposaurid found in Russia. More recent studies of therocephalians have found scaloposaurids like Scalopodon to be juvenile forms of larger therocephalians and do not consider Scaloposauridae to be a valid group. Scalopodon and most other scaloposaurids are now classified as basal members of Baurioidea.
Lycideops is an extinct genus of therocephalians from the Late Permian of South Africa. The type species is Lycideops longiceps, named in 1931 by South African paleontologist Robert Broom. Fossils of Lycideops come from the Dicynodon Assemblage Zone of the Beaufort Group. Lycideops is a member of the family Lycideopidae. Like other lycideopids, Lycideops has a long snout.
Yikezhaogia is an extinct genus of therocephalian therapsids from the Early Triassic of Mongolia. It is known from a single fragmentary skull and associated postcranial bones representing the species Yikezhaogia megafenestrala. It is identifiable as a therocephalian by its thin postorbital bar behind the eye socket, its elongated temporal opening behind the bar, and a thin lower jaw with a low coronoid process. Large tooth sockets in the upper jaw indicate that Yikezhaogia had large caniniform teeth. The teeth of the lower jaw are blunt-tipped and cylindrical. Although its exact position among therocephalians is uncertain, Yikezhaogia is probably a basal member of the group Baurioidea.
Simorhinella is an extinct genus of therocephalian therapsids from the Late Permian of South Africa. It is known from a single species, Simorhinella baini, named by South African paleontologist Robert Broom in 1915. Broom named it on the basis of a single fossil collected by the British Museum of Natural History in 1878 that included the skull and jaws forward from the eye sockets. The skull is unusual in that it has an extremely short and deep snout, unlike the longer and lower snouts of most other therocephalians. Because of the skull's distinctiveness, the classification of Simorhinella within Therocephalia is uncertain. However, a 2014 study proposed that it was closely related to the basal therocephalian Lycosuchus, placing it in the family Lycosuchidae.
Kembawacela is an extinct genus of cistecephalid dicynodont from the Late Permian Madumabisa Mudstone Formation of Zambia. It contains the type and only species, Kembawacela kitchingi. Like other cistecephalids, Kembawacela was specialised for a fossorial, burrowing lifestyle similar to modern day moles. It is unique amongst cistecephalids for the presence of a pair of tusks in the upper jaw, characteristic of many other dicynodonts but lost in other cistecephalids. It is likely that Kembawacela was a locally endemic species of cistecephalid in the Luangwa Basin of Zambia.
