In the geologic timescale, the Wuchiapingian or Wujiapingian is an age or stage of the Permian. It is also the lower or earlier of two subdivisions of the Lopingian Epoch or Series. The Wuchiapingian spans the time between 259.51 and 254.14 million years ago (Ma). It was preceded by the Capitanian and followed by the Changhsingian.
Therocephalia is an extinct clade of eutheriodont therapsids from the Permian and Triassic. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their teeth, suggest that they were carnivores. Like other non-mammalian synapsids, therocephalians were once described as "mammal-like reptiles". Therocephalia is the group most closely related to the cynodonts, which gave rise to the mammals. This relationship takes evidence in a variety of skeletal features.
Euchambersia is an extinct genus of therocephalian therapsids that lived during the Late Permian in what is now South Africa and China. The genus contains two species. The type species E. mirabilis was named by paleontologist Robert Broom in 1931 from a skull missing the lower jaw. A second skull, belonging to a probably immature individual, was later described. In 2022, a second species, E. liuyudongi, was named by Jun Liu and Fernando Abdala from a well-preserved skull. It is a member of the family Akidnognathidae, which historically has also been referred by as the synonymous Euchambersiidae.
Moschorhinus is an extinct genus of therocephalian in the family Akidnognathidae with only one species: M. kitchingi. It was a carnivorous synapsid which has been found in the Late Permian to Early Triassic of the South African Karoo Supergroup. It was a large carnivore, reaching 1.5 m (4.9 ft) in total body length with the largest skull comparable to that of a lion in size. It had a broad, blunt snout which bore long, straight canines. It appears to have replaced the gorgonopsids ecologically, and hunted much like a big cat. While most abundant in the Late Permian, it survived a little after the Permian Extinction, though these Triassic individuals had stunted growth.
Moschowhaitsia is an extinct genus of therocephalian therapsids from the Late Permian (Guadalupian) of Russia and China. The type species, Moschowhaitsia vjuschkovi, was discovered in the Changxingian-aged Archosaurus Assemblage Zone of Russia and named in 1963 by Russian palaeontologist Leonid Petrovich Tatarinov. A second species was discovered in Jingtai County of Gansu, China in 2020 and named as M. lidaqingi in 2023 by Jun Liu and Fernando Abdala, the first whaitsiid therocephalian to be discovered in China. It was among the larger carnivores in the faunal assemblages it occurred in, with a skull-length of up to 25 centimetres (9.8 in) in M. vjuschkovi and an even larger estimated 35 centimetres (14 in) for M. lidaqingi, one of the largest therocephalian skulls reported. The genus name Moschowhaitsia alludes to two other therocephalians, Moschorhinus and Whaitsia, due to the structure of its palate combining physical features of both these genera.
Chlynovia is an extinct genus of therocephalian therapsids from the Late Permian of Russia. The type species is C. serridentatus, named in 2000. Chlynovia was originally classified within Scaloposauria, a group of therocephalians characterized by their small size and lightly built skulls. Scaloposaurians are no longer recognized as a true grouping, but instead represent the juvenile forms of many types of therocephalians. Chlynovia was placed in the family Perplexisauridae along with Perplexisaurus, but both therocephalians are now placed in the family Ictidosuchidae.
Scalopodontes is an extinct genus of therocephalian which existed in Russia during the Late Permian period. The type species is Scalopodontes kotelnichi. Fossils have been found in the Sokolki Assemblage Zone of the Urpalov Formation.
Daqingshanodon is an extinct genus of dicynodont therapsid from the Late Permian of Inner Mongolia, China. The type species D. limbus was described in 1989 from a single skull found in the Naobaogou Formation. Daqingshanodon belongs to a group of dicynodonts called cryptodonts. It is the smallest known cryptodont, and the only one known from China. Like other cryptodonts, it has a pair of rounded nasal bosses above its nostrils and a ridge of bone on the upper jaw called the postcaniniform process. Daqingshanodon has a pair of elongated, recurved tusks extending from its beak-like snout. It is distinguished from other dicynodonts by the presence of a distinct ridge running along the side of the skull from below the eye socket to the area around the tusks. The skull of Daqingshanodon is less than 10 centimetres (3.9 in) long, yet this specimen is thought to have been an adult on the basis of its well-developed nasal bosses.
Ordosiodon is an extinct genus of therocephalian therapsids from the Early Triassic of China. It includes two species, O. lincheyuensis and O. youngi.
Jimusaria is an extinct genus of dicynodont therapsid from the Late Permian (Changhsingian) of China. The type species J. sinkianensis from the Guodikeng Formation in Xinjiang, was originally named as a species of Dicynodon, the first from Asia, but was given its own genus in 1963 before being sunk back into Dicynodon in 1988. The genus was resurrected in 2011 by palaeontologist Christian Kammerer in a taxonomic revision of the genus Dicynodon. Jimusaria was a mid-sized dicynodont, and was similar in appearance to the South African Dicynodon, but differed from it in features such as its narrower snout. A second species, Jimusaria monanensis was described from the Naobaogou Formation of northern China in 2023.
Turfanodon is an extinct genus of dicynodont therapsid from the Late Permian Sunan, Guodikeng, and Naobaogou Formations of China. The holotype of T. bogdaensis was discovered between 1963-1964 and was originally named in 1973 by A. Sun with the type species Turfanodon bogdaensis, Turfanodon was reclassified as a junior synonym of the related Dicynodon in 1988 by G. M. King. T. bogdaensis remained a species of Dicynodon for over two decades before the genus was reinstated in 2011 in a revision of the taxonomy of Dicynodon by palaeontologist Christian Kammerer. A second species from Inner Mongolia, T. jiufengensis, was named in 2021 by palaeontologist Jun Liu from a nearly complete skeleton and other referred bones. Turfanodon was a relatively large dicynodont, and similar in appearance to the related Daptocephalus from South Africa.
Hazhenia is an extinct genus of therocephalian therapsids from the Early Triassic of China, of which Hazhenia concava is the only species. Hazhenia was named in 1981 from the Heshanggou Formation in the Ordos Desert of Inner Mongolia. It lived during the Olenekian Age of the Early Triassic, about 247 million years ago. Hazhenia belongs to a group of therocephalians called Baurioidea and possesses many mammal-like features such as cusped teeth and a secondary palate, both of which evolved independently in baurioids. Within Baurioidea it is most closely related to the genus Ordosiodon, which is also known from Inner Mongolia but comes from the slightly younger Ermaying Formation. Both genera were once placed in the family Ordosiidae, but as the name is preoccupied by a family of Cambrian trilobites, it is no longer valid.
Ichibengops is an extinct genus of therocephalian therapsids known from the type species Ichibengops munyamadziensis, which lived in what is now Zambia during the Late Permian. Ichibengops was named in 2015 on the basis of fossils found in the Wuchiapingian-age Madumabisa Mudstone Formation in the Luangwa Basin. Therocephalians have been known from the Luangwa Basin for decades, yet Ichibengops was the first endemic Zambian therocephalian to have been described in detail. Phylogenetic analysis indicates that it is a basal member of the clade Eutherocephalia, lying just outside a clade containing hofmeyriids, whaitsiids, and baurioids. Ichibengops is the sister taxon of the Russian therocephalian Chthonosaurus; together they form one of several known African-Russian sister taxon pairs of eutherocephalians, which indicate that eutherocephalians could freely disperse across most of Pangea during the Late Permian. Like the fellow therocephalian Euchambersia, Ichibengops might have had venom glands, as evidenced by grooves above its teeth.
Dalongkoua fuae is a Chinese whaitsiid therocephalian of the Late Permian.
Ericiolacertidae is a family of therocephalian therapsids. The family living in the earliest Triassic after the Permian-Triassic extinction event. Genera are: Ericiolacerta found in the Fremouw Formation of Antarctica and the Normandien Formation of South Africa, Pedaeosaurus found in the Fremouw Formation of Antarctica and Silphedosuchus, from the Petropavlovka Formation of Russia, east of the Urals. The family is composed of carnivores.
Gorynychus is a genus of therocephalian from the mid-Permian from Kotelnich, Russia. The genus contains two species, G. masyutinae and G. sundyrensis. It was named after the three-headed dragon Zmey Gorynych from Russian mythology.
The Naobaogou Formation is a geological formation in the Daqing Mountains of China. It is likely of Lopingian age. It consists of three rhythms of sediment, labelled members I-III primarily of purple siltstone, but each with a thick basal conglomerate bed. It is notable for its fossil content, producing one of the most diverse Late Permian vertebrate faunas outside Russia and South Africa.
Caodeyao is a genus of therocephalian that lived in what is now China during the late Permian. It was found in the Naobaogou Formation. It contains one species, Caodeyao liuyufengi, named in 2020 by Jun Liu and Fernando Abdala.
The Sunjiagou Formation is a geological formation in Shanxi, China. It is of Lopingian age. The lower and middle parts of the formation consists of intensely bioturbated fine grained sandstones and thinly interbedded mudstones, deposited in a shallow-shore lake depositional environment, while the upper part consists of fine grained sandstone, siltstone and mudstone. Alongside the Naobaogou Formation, it has provided an important vertebrate fauna.
Jiufengia is an extinct genus of therocephalian in the family Akidnognathidae. It is known from a single species, Jiufengia jiai, from the Late Permian Naobaogou Formation in China.
