Clouded yellows | |
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Colias hyale type species for the genus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Lepidoptera |
Family: | Pieridae |
Tribe: | Coliadini |
Genus: | Colias Fabricius, 1807 |
Type species | |
Papilio hyale | |
Synonyms | |
Colias is a genus of butterflies in the family Pieridae. They are often called clouded yellows in the Palearctic and sulphurs (a name also used for other coliadine genera) in North America. The closest living relative is the genus Zerene , [4] which is sometimes included in Colias.
This genus occurs throughout the Holarctic, including the arctic regions. They are also found in South America, Africa, China and India. Their caterpillars feed on certain Fabaceae, for example vetches (Vicia). While most are thus beneficial by keeping weeds at bay, some occasionally become nuisance pests on crops like alfalfa. In some species, the wings of males have brilliant ultraviolet reflection, while those of females do not. [5] [6] Adults of both sexes have various colour forms.
Most if not all species of this genus, as usual for Coliadinae, do not sequester toxins or other noxious compounds from their food plants. They are therefore a well-loved prey item of insectivores as compared to Pieris of the related Pierinae. They make up this disadvantage by being more nimble and better able to evade attacks by would-be predators. [7]
Notable lepidopterologists who did many studies on this genus included Julius Röber, J. Malcolm Fawcett, George B. Johnson and Henry Rowland-Brown.
Hybridization runs rampant in these polytypic and clinal [8] butterflies, confounding molecular phylogenetics studies. In general, cladistic analyses of only one type of data (particularly mtDNA sequences) cannot be considered reliable. Regardless, the evolutionary distance within some "species" is so large that cryptic speciation rather than (or in addition to) interbreeding seems to be the cause. For example, the Beringian populations traditionally assigned to the northern clouded yellow (C. hecla) could warrant recognition as a species; hybridization between North American and Asian populations seems to have played a role in their evolution, but as a whole they appear to be a rather old and distinct lineage. [9]
Listed alphabetically: [10]
Colias are usually some shade of yellow, orange or white. Their uppersides feature black borders (usually solid in males, often with pale spots in females). They always perch with wings closed, but upperside pattern may be seen faintly through the wing, or glimpsed in flight. [11]
The Pieridae are a large family of butterflies with about 76 genera containing about 1,100 species, mostly from tropical Africa and tropical Asia with some varieties in the more northern regions of North America and Eurasia. Most pierid butterflies are white, yellow, or orange in coloration, often with black spots. The pigments that give the distinct coloring to these butterflies are derived from waste products in the body and are a characteristic of this family. The family was created by William John Swainson in 1820.
Colias croceus, clouded yellow, is a small butterfly of the family Pieridae, the yellows and whites.
Colias hyale, the pale clouded yellow, is a butterfly of the family Pieridae, which is found in most of Europe and large parts of the Palearctic. It is a rare migrant to the British Isles and Scandinavia. The adult wingspan is 52–62 millimetres (2.0–2.4 in).
Colias alfacariensis, Berger's clouded yellow, is a butterfly of the family Pieridae. It was separated from the pale clouded yellow, C. hyale, in 1905. Berger's clouded yellow is a Palearctic species (South and Central Europe, South Russia, Russian Far East, Siberia Central Asia and temperate China also Asia Minor, Caucasus and Transcaucasia.
Colias palaeno, known by the common names moorland clouded yellow, palaeno sulphur, and pale Arctic clouded yellow, is a butterfly in the family Pieridae.
Oeneis is a butterfly genus of the Satyrinae. All but one of its members are Arctic, sub-Arctic or high-altitude alpine in distribution. Some of the members of the genus are among the butterflies that can get along in the harshest climates of any butterflies. Four species in Europe, more are found in Arctic Russia, Siberia, Mongolia, Arctic North America and the Rocky Mountains. Curiously, there are no observations from Greenland. The development of most species takes two years.
Colias philodice, the common sulphur or clouded sulphur, is a North American butterfly in the family Pieridae, subfamily Coliadinae.
Colias aurorina, the Greek clouded butterfly or dawn clouded yellow, is a butterfly in the family Pieridae. It is found in Greece, the Near East, the Caucasus, and in Iran and Turkmenistan.
Colias phicomone, the mountain clouded yellow, is a butterfly in the family Pieridae. It is found in the Cantabrian Mountains, the Pyrenees, the Carpathian Mountains and the Alps. It flies at altitudes of 900 to 2800 meters.
Colias nastes, the Labrador sulphur, is a butterfly in the family Pieridae. In Europe, it is found in the north of Norway and Sweden and on rare occasions in northern Finland. It is also found in North America, specifically in Alaska, Canada, and the Rocky Mountains, Washington, Montana and on Greenland. In Asia, it is found in the Altai Mountains, the border regions of Russia, China, Mongolia, Kazakhstan, the Sayan Mountains, the north of Siberia, and in the Chukotka Autonomous Okrug.
Colias tyche, the Booth's sulphur or pale Arctic clouded yellow, is a butterfly in the family Pieridae. It is found from Baffin Island west along the Hudson Bay and arctic coasts of the Nunavut and Northwest Territories mainland and the southern tier of Arctic Islands to northern Yukon, Alaska, and Eurasia.
Colias wiskotti is a butterfly in the family Pieridae. It is found in Turkestan, Uzbekistan, Afghanistan, Tajikistan, and Kashmir. The habitat consists of xerophytic mountains.
Colias alpherakii is a butterfly in the family Pieridae found in Central Asia.
Colias felderi is a butterfly in the family Pieridae. It is found in Tibet and China.
Colias hyperborea is a butterfly in the family Pieridae. It is found in the East Palearctic.
Colias "Le genre Colias" provides distribution information in French.