Dynastes | |
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Dynastes grantii | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Coleoptera |
Family: | Scarabaeidae |
Tribe: | Dynastini |
Genus: | Dynastes MacLeay, 1819 |
Synonyms | |
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Dynastes is a genus of large beetles belonging to the family Scarabaeidae. They occur in the Nearctic realm and in the Neotropical realm, from the United States to Brazil; [1] [2] four North American species (including Mexico), three with distributions extending from Central America either north or south, and two species endemic to South America. [2]
Males of Dynastes bear two long horns, one on the head, and the other on the pronotum, forming a "plier"; the pronotal horn has reddish setae on its underside. This pronotal horn is absent in females. [3] Some species have an iridescent colouration to their elytra. [4] Certain species of the genus Dynastes also have the ability to change colour. [5] Specific species have been noted to occur with either black or yellowish to khaki green elytra. [5] This variation in colour is due to a spongy layer below the transparent cuticle; [5] this spongy layer is a network of filamentous strands made up of three-dimensional photonic crystals lying parallel to the cuticle surface. [6] When the cuticle is filled with gas this layer can show through, presenting the yellow to khaki green colour, but when filled with fluid the cuticle appears black. [5] This is due to the change in refraction index allowing us to see the difference in colours. [6] This system is known as a hygrochromic effect. [4] Female Dynastes can change colour but not as completely as males, which is not yet explained as the mechanism for the colour change is still not completely understood. [5] What is known is that changes in humidity affect the levels of moisture in the cuticle which leads to a change in colour in most cases. [5] Since the change is due to humidity it is a reversible process, however, it has been observed that after multiple colour changes or high stress the beetles will maintain some dark spots on their cuticle. [4] Some hypotheses for why this colour change occurs at all are the ability to blend with surroundings depending on the time of day (black for nighttime and yellow for daytime) to best avoid their main predator, the tropical screech owl (Megascops choliba). [5] Another theory has to do with thermoregulation in the sense that a black beetle heats up faster than yellow and then once they have warmed up theoretically there will be less moisture in the cuticle which leads to changing to a colour which does not heat as quickly so they won't overheat. [5]
There are eight species currently recognized in the genus, not counting putative subspecies of D. hercules (Hercules beetle):
Male | Female | Larvae | Scientific name | Common Name | Distribution |
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Dynastes grantii Horn, 1870 | Western Hercules beetle or Grants' Hercules beetle | southwestern United States (Arizona, New Mexico, Utah), northern Mexico | |||
Dynastes hercules (Linnaeus, 1758) | Hercules beetle | southern Mexico, Central and South America, Lesser Antilles | |||
Dynastes hyllus Chevrolat, 1843 | Mexico, Belize, El Salvador, Honduras, Guatemala, Nicaragua. Male: 35–70 mm (1.4–2.8 in); female: 30–45 mm (1.2–1.8 in) | ||||
Dynastes maya Hardy, 2003 | Maya white beetle [7] | southern Mexico, Guatemala, Honduras. Male: 50–90 mm (2.0–3.5 in); female: 40–50 mm (1.6–2.0 in) | |||
Dynastes moroni Nagai, 2005 | Sierra de los Tuxtlas in Mexico | ||||
Dynastes neptunus (Quensel in Schönherr, 1805 | Neptune beetle | northwestern South America (Colombia, Ecuador, Peru, Venezuela) | |||
Dynastes satanas Moser, 1909 | Satanas beetle | Bolivia. Male: 50–115 mm (2.0–4.5 in); female: 30–55 mm (1.2–2.2 in) | |||
Dynastes tityus (Linnaeus, 1763) | Unicorn beetle or Eastern Hercules beetle | eastern United States (New York and Florida west to Illinois and Texas) [8] | |||
Although there are numerous species under the genus Dynastes, some are able to produce viable offspring with one another. [2] This has been observed in captivity, but it is unclear if wild beetles will engage in acts of hybridization. [2] Certain species such as D. grantii and D. hyllus are believed to be sister species, while D. tityus is thought to be a sister taxon to the Central American "white Hercules" lineage. [2] The intermediate species that bridges the "white Hercules" and the "giant Hercules" lineages is thought to be D. maya. [2]
The larval stage of Dynastes hercules will last one to two years, with the larva growing up to 4.5 inches (11 cm) in length and weighing more than 100 g (3.5 oz). Much of the life of the larva is spent tunneling through rotting wood. After the larval period, transformation into a pupa, and moulting, the beetle then emerges as an adult. Adults of most species can live from two to ten months [9] and some can even live one or two years. Eastern Hercules beetles, D. tityus, can live six to twenty-three months in captivity with a hibernation period. [9] Western Hercules beetles, D. grantii, tend to have a shorter adult lifespan in the wild (two to four months), but in captivity they live for about the same amount of time as the eastern species. [9] It has also been noted that captive longevity is possible without a hibernation period. [9]
The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change in recent years. Several subfamilies have been elevated to family rank, and some reduced to lower ranks. The subfamilies listed in this article are in accordance with those in Catalog of Life (2023).
Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Its constituent families are also undergoing revision presently, and the family list below is only preliminary. This superfamily includes some of the largest beetles extant today, including rhinoceros beetles, (Dynastinae), the Hercules beetle and Goliath beetles.
The Hercules beetle is a species of rhinoceros beetle native to the rainforests of southern Mexico, Central America, South America, and the Lesser Antilles. It is the longest extant species of beetle in the world, and is also one of the largest flying insects in the world.
Dynastinae or rhinoceros beetles are a subfamily of the scarab beetle family (Scarabaeidae). Other common names – some for particular groups of rhinoceros beetles – include Hercules beetles, unicorn beetles or horn beetles. Over 1500 species and 225 genera of rhinoceros beetles are known.
Dynastes tityus, the eastern Hercules beetle, is a species of rhinoceros beetle native to the Eastern United States. The adult's elytra are green, gray or tan, with black markings, and the whole animal, including the male's horns, may reach 60 mm (2.4 in) in length. The larvae feed on decaying wood from various trees.
Megaceras briansaltini, named in honor of Brian Saltin, son of entomologist Jochen-P. Saltin, is a Peruvian rhinoceros beetle with a horn surprisingly similar to that of the character Dim from Pixar's animated film, A Bug's Life, previously unseen in nature. It remains unclear whether this is a natural feature or an uncommon mutation, as only one specimen has been found.
Dynastes hyllus is a large scarab beetle species that ranges from Mexico to Guatemala. Its larvae have been found to associate with the logs of Persea americana.
Dynastes satanas, the Satanas beetle, is a species of beetle belonging to the family Scarabaeidae. The name is sometimes misspelled as "satanus".
The western Hercules beetle is a species of rhinoceros beetle that lives in Arizona, New Mexico and Utah in the United States and in parts of northern Mexico. This species is known for its grayish-white elytra, large size, and characteristic horn of the adult males.
The Moellenkampi beetle, is one of four large species of rhinoceros beetles from Southeast Asia belonging to the genus Chalcosoma. The Moellenkampi beetles belong to the beetle family Scarabaeidae along with other rhinoceros beetles. They are characterized by having two large, forward-projecting horns on the pronotum or thorax, and another large, forward and upward-projecting horn on the head. They also have a distinct metallic lustre, which is the reason behind their genus name, which derives from Greek, chalko-, and is the combining form of chalkós, meaning copper.
Chalcosoma chiron is a species of beetle in the family Scarabaeidae. This species can be found from Malaysia south into Indonesia and Thailand in East region. It was formerly known as Chalcosoma caucasus, a name which is a junior synonym and not valid.
Inca clathratus is a species of flower chafer in the family Scarabaeidae, indigenous to Central and South America. These beetles can be seen surrounding the sap flows from injured or diseased trees anytime in the day and sometimes at night. They mostly feed at the sap flows of the trees, including avocado and mango trees. Like other beetles they also feed off of rotting fruit. There are a few different subspecies of Inca clathratus including: Inca clathratus sommeri, Inca clathratus clathratus, and Inca clathratus quesneli. The species name has been frequently misspelled as "clathrata" but the genus name Inca is masculine.
Dynastes moroni is a large scarab beetle endemic to the Sierra de los Tuxtlas region in Mexico.
Margarinotus ephemeralis is a species of clown beetle in the family Histeridae. It is currently only known from southern California.
Caccobius unicornis, is a species of dung beetle found in many Asian and South East Asian countries such as: India, Sri Lanka, China, Java, Borneo, Taiwan, Korea, Japan, Myanmar, North Vietnam, Philippines, Sumatra, Java, Borneo, Malaysia, Thailand, and Indochina.
Pseudaspidimerus mauliki, is a species of lady beetle native to India, Sri Lanka, Bangladesh, Thailand, and Java.
Telegeusinae is a small subfamily of beetles in the family Omethidae recognizable by enlarged palpi found in males. Though relatively rare, males are sometimes found in large numbers in black light traps. Females are not known in this group, but it is theorized that females are larviform as found in many closely related taxa.
Dynastes neptunus is a beetle in the family Scarabaeidae. D. neptunus is very simular to the Hercules beetle but differs from it in slightly larger size and more of a slender and curved horns.
Oryctoantiquus is a genus of middle Eocene beetles in the Dynastinae clade.