Halitherium | |
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Halitherium schinzi skeleton, Muséum national d'Histoire naturelle, Paris | |
Life restoration of H. schinzii | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Sirenia |
Family: | Dugongidae |
Subfamily: | † Halitheriinae |
Genus: | † Halitherium Kaup, 1838 |
Species | |
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Halitherium is an extinct dugongid sea cow that arose in the late Eocene, then became extinct during the early Oligocene. Its fossils are common in European shales. Inside its flippers were finger bones that did not stick out. Halitherium also had the remnants of back legs, which did not show externally. However, it did have a basic femur, joined to a reduced pelvis. Halitherium also had elongated ribs, presumably to increase lung capacity to provide fine control of buoyancy. A 2014 review presented the opinion that the genus is dubious.
Halitherium is the type genus of the subfamily Halitheriinae, which includes the well-known genera Eosiren and Eotheroides and lived from the Eocene to the Oligocene. [1]
The genus Halitherium has had a confusing nomenclatural history. [2] It was originally coined by Johann Jakob Kaup on the basis of a premolar from the early Oligocene (Rupelian) of southern Germany, but Kaup himself mistakenly stated that the premolar, in his opinion, gehort zu Hippopotamus dubius Cuv., unaware that H. dubius is actually a junior synonym of the primitive sirenian Protosiren minima , while simultaneously coining the genus and species name Pugmeodon schinzii for the same specimen. [3] For his part, the renowned German paleontologist Christian Erich Hermann von Meyer included the type specimen of Halitherium schinzii in his composite species Halianassa studeri, [4] whose hypodigm also included the type specimens of Metaxytherium medium and Protosiren minima as well as a Miocene-age maxilla and a skeleton from the molasse basin in Switzerland. [5] [6]
Later, Kaup synonymized Pugmeodon with Halitherium creating the new combination Halitherium schinzii, and the name Halitherium became universally accepted for the early Oligocene halitheriine material from Europe. [7] Because Halitherium was originally based on a misidentified type species and due to the widespread use of Halitherium, the sirenian specialist Daryl Domning petitioned the ICZN to designate Pugmeodon schinzii as the type species of Halitherium, and the proposal was approved by the Commission in 1989, effectively making Pugmeodon a junior objective synonym of Halitherium in line with the current concept of Halitherium introduced by Kaup himself. [8] [9]
Voss (2013, 2014) dismisses Halitherium as a nomen dubium by virtue of being based on non-diagnostic remains. Voss based the opinion on the type species, H. schinzii, being nomen dubium , with its holotype fossil, an isolated molar, having no diagnostic value. [10] and a 2017 study found specimens traditionally assigned to Halitherium schinzii to be two separate species, one of which takes the name Halitherium bronni Krauss, 1858. Because Halitherium is dubious, H. bronni has been re-assigned to Kaupitherium . [10] [11] The species Halitherium alleni Simpson, 1932, described by Simpson (1932) from skull caps in Oligocene deposits in Puerto Rico, [12] is basal to the two Kaupitherium species. [13]
The Sirenia, commonly referred to as sea cows or sirenians, are an order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The extant Sirenia comprise two distinct families: Dugongidae and Trichechidae with a total of four species. The Protosirenidae and Prorastomidae families are extinct. Sirenians are classified in the clade Paenungulata, alongside the elephants and the hyraxes, and evolved in the Eocene 50 million years ago (mya). The Dugongidae diverged from the Trichechidae in the late Eocene or early Oligocene.
Dugongidae is a family in the order of Sirenia. The family has one surviving species, the dugong, one recently extinct species, Steller's sea cow, and a number of extinct genera known from fossil records.
Trichechidae is a family of sirenians that includes all living manatees and several extinct genera.
The Desmostylia are an extinct order of aquatic mammals native to the North Pacific from the early Oligocene (Rupelian) to the late Miocene (Tortonian). Desmostylians are the only known extinct order of marine mammals.
Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, prompting attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.
Paleoparadoxia is a genus of large, herbivorous aquatic mammals that inhabited the northern Pacific coastal region during the Miocene epoch. It ranged from the waters of Japan, to Alaska in the north, and down to Baja California, Mexico.
Behemotops is an extinct genus of herbivorous marine mammal. It lived from the Early Oligocene (Rupelian) through the Late Oligocene, existing for approximately 10.9 million years. It is the most primitive known desmostylian, believed to be close to the ancestry of all other desmostylians.
Eotheroides is an extinct genus of Eocene sirenian. It is an early member of the family Dugongidae, which includes the extant dugong. Fossils have been found from Egypt, India, and Madagascar. Eotheroides was first described by Richard Owen in 1875 under the name Eotherium, which was replaced by the current name in 1899.
Metaxytherium is an extinct genus of dugong that lived from the Oligocene until the end of the Pliocene. Fossil remains have been found in Africa, Europe, North America and South America. Generally marine seagrass specialists, they inhabited the warm and shallow waters of the Paratethys, Mediterranean, Caribbean Sea and Pacific coastline. American species of Metaxytherium are considered to be ancestral to the North Pacific family Hydrodamalinae, which includes the giant Steller's sea cow.
Protosiren is an extinct early genus of the order Sirenia. Protosiren existed throughout the Lutetian to Priabonian stages of the Middle Eocene. Fossils have been found in the far-flung locations like the United States, Africa (Egypt), Europe and Asia.
Eosiren is an extinct genus of sea cow that lived during the Late Eocene to Early Oligocene (Rupelian). Several fossils have been found in Egypt. It seems like the species E. abeli were contemporaneous with Protosiren and Eotheroides. like them, Eosiren closely resembled modern sirenians. It differs from them by having somewhat larger innominates and possess thigh bones.
Dusisiren is an extinct genus of dugong related to the Steller's sea cow that lived in the North Pacific during the Neogene.
Miosiren is an extinct genus of manatee from the Early Miocene of southeastern England (Suffolk) and Antwerp, Belgium.
Anomotherium is an extinct genus of manatee that lived in the shallow seas of what is now northern Germany. Its closest relative is Miosiren. Fossils of the genus have been found in the Bohlen and Doberg Formations of Germany.
Potamosiren is an extinct genus of manatee from the Middle Miocene (Laventan) Honda Group of Colombia.
Lentiarenium was an early sea cow from the Late Oligocene (Chattian) Linz-Melk Formation of Austria. Known since the mid 19th century, Lentiarenium was long considered to be a species of Halitherium until a 2016 analysis showed it to be distinct.
Italosiren is an extinct genus of early dugong from the Early Miocene (Aquitanian) Libano Formation in Northern Italy.
Kaupitherium is an extinct dugongid sea cow that lived during the Oligocene. Fossils of the genus have been found in the Alzey Formation of Germany. Inside its flippers were finger bones that did not stick out. Kaupitherium also had the residues of back legs, which did not show externally. However, it did have a basic femur, joined to a reduced pelvis. Kaupitherium also had elongated ribs, presumably to increase lung capacity to provide fine control of buoyancy.
Crenatosiren is an extinct genus of dugongid sirenian known from the late Oligocene (Chattian) of Florida, North Carolina, and South Carolina. The type and only known species is Crenatosiren olseni.
Stegosiren was an early sea cow from the Middle Oligocene of South Carolina, US. It shows a stage of halitheriine evolution more derived than that of the Old World early Oligocene Eosiren imenti and Halitherium schinzii.