Eotheroides

Last updated

Eotheroides
Temporal range: Middle - Late Eocene, 48.6–33.9  Ma
Eotheroides sp.JPG
Eotheroides sp. in Artis, Amsterdam.
Scientific classification
Kingdom:
Phylum:
Class:
Order:
Family:
Genus:
Eotheroides

Trouessart, 1905
Species
  • E. aegyptiacum(Owen, 1875) (type)
  • E. babiaeBajpai et al., 2006
  • E. clavigerumZalmout and Gingerich, 2012
  • E. lambondranoSamonds et al., 2009
  • E. majusZdansky, 1938
  • E. sandersiZalmout and Gingerich, 2012
  • E. waghapadarensisDas and Basu, 1994

Eotheroides is an extinct genus of Eocene sirenian. It is an early member of the family Dugongidae, which includes the extant dugong. Fossils have been found from Egypt, India, and Madagascar. Eotheroides was first described by Richard Owen in 1875 under the name Eotherium, which was replaced by the current name in 1899.

Contents

Based on endocranial casts, Eotheroiodes had a smaller endocranial volume than other contemporaneous sirenians such as Protosiren . Unlike extant sirenians, Eotheroides possesses a tentorium cerebelli, which appears as a distinct transverse groove on the skull. [1]

Species

Drawing of skeleton from the species E. sandersi Eotheroides remains.png
Drawing of skeleton from the species E. sandersi

The type species, E. aegyptiacum, is known from the Lutetian Mokattam Limestone of Cairo, Egypt. [1] Another species, E. lambondrano, was recently[ when? ] named on the basis of material found from Middle Eocene nearshore marine deposits in the Mahajanga Basin of Madagascar. The species was named after the Malagasy word for dugong, which translates as "water bushpig". It is known from a nearly complete skull and fragments of pachyosteosclerotic (thick) ribs. Based on its age and morphology, E. lambondrano may be ancestral to E. aegyptiacum. [2] Described in 2009, E. lambondrano is the first pre-Pleistocene Cenozoic mammal named from Madagascar, being known from an 80-million-year gap in the island's fossil record. [3]

Several other species have been named, including E. babiae from India, [4] E. majus, E. clavigerum and E. sandersi from Egypt, [5] [6] and E. waghapadarensis, also from India. [7] E. majus was based on a single upper molar, which was never catalogued and is now lost; [6] it has been considered a senior subjective synonym of E. aegyptiacum. [8]

Paleobiology

Drawing of swimming Eotheroides sandersi (green), compared to three other Eocene sirenian taxa. Eocene sirenians size comparison.png
Drawing of swimming Eotheroides sandersi (green), compared to three other Eocene sirenian taxa.

The teeth of Eotheroides were relatively unspecialized compared to those of extant sirenians, which are reduced as an adaptation for feeding on sea grass. The upper molars of E. lambondrano are considerably longer and wider than those of E. aegyptiacum, suggesting that they were less specialized. [1] Eotheroides is likely to have been one of the first fully aquatic sea cows. Eotheroides lambondrano was found in association with the remains of other sea cows, crocodilians, and sea turtles, which suggests that the locality is representative of a coastal or estuarine environment. [3]

Related Research Articles

<span class="mw-page-title-main">Sirenia</span> Order of aquatic herbivorous mammals

The Sirenia, commonly referred to as sea cows or sirenians, are an order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The extant Sirenia comprise two distinct families: Dugongidae and Trichechidae with a total of four species. The Protosirenidae and Prorastomidae families are extinct. Sirenians are classified in the clade Paenungulata, alongside the elephants and the hyraxes, and evolved in the Eocene 50 million years ago (mya). The Dugongidae diverged from the Trichechidae in the late Eocene or early Oligocene.

<span class="mw-page-title-main">Dugongidae</span> Family of mammals

Dugongidae is a family in the order of Sirenia. The family has one surviving species, the dugong, one recently extinct species, Steller's sea cow, and a number of extinct genera known from fossil records.

<i>Hydrodamalis</i> Extinct family of mammals

Hydrodamalis is a genus of extinct herbivorous sirenian marine mammals, and included the Steller's sea cow, the Cuesta sea cow, and the Takikawa sea cow. The fossil genus Dusisiren is regarded as the sister taxon of Hydrodamalis: together, the two genera form the dugong subfamily Hydrodamalinae. They were the largest member of the order Sirenia, whose only extant members are the dugong and the manatees. They reached up to 9 metres (30 ft) in length, making the Steller's sea cow among the largest mammals other than whales to have existed in the Holocene epoch. Steller's sea cow was first described by Georg Wilhelm Steller,

<i>Basilosaurus</i> Prehistoric cetacean genus from the Late Eocene epoch

Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, which prompted attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.

<i>Dorudon</i> Genus of ancient whales

Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring 5 metres (16 ft) long and weighing 1–2.2 metric tons. Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand and Western Sahara.

<span class="mw-page-title-main">Wadi al Hitan</span> Paleontological site in the Faiyum Governorate of Egypt

Wādī al-Ḥītān is a paleontological site in the Faiyum Governorate of Egypt, some 150 kilometres (93 mi) south-west of Cairo. It was designated a UNESCO World Heritage Site in July 2005 for its hundreds of fossils of some of the earliest forms of whale, the archaeoceti. The site reveals evidence for the explanation of one of the greatest mysteries of the evolution of whales: the emergence of the whale as an ocean-going mammal from a previous life as a land-based animal.

<span class="mw-page-title-main">Archaeoceti</span> Paraphyletic group of primitive cetaceans from Early Eocene to Late Oligocene

Archaeoceti, or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene. Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia 53 to 45 mya, resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation 36 to 35 mya.

<span class="mw-page-title-main">Evolution of sirenians</span> Development from a Tethytherian ancestor and radiation of species

Sirenia is the order of placental mammals which comprises modern "sea cows" and their extinct relatives. They are the only extant herbivorous marine mammals and the only group of herbivorous mammals to have become completely aquatic. Sirenians are thought to have a 50-million-year-old fossil record. They attained modest diversity during the Oligocene and Miocene, but have since declined as a result of climatic cooling, oceanographic changes, and human interference. Two genera and four species are extant: Trichechus, which includes the three species of manatee that live along the Atlantic coasts and in rivers and coastlines of the Americas and western Africa, and Dugong, which is found in the Indian and Pacific oceans.

Protosiren is an extinct early genus of the order Sirenia. Protosiren existed throughout the Lutetian to Priabonian stages of the Middle Eocene. Fossils have been found in the far-flung locations like the United States, Africa (Egypt), Europe and Asia.

<span class="mw-page-title-main">Remingtonocetidae</span> Family of mammals

Remingtonocetidae is a diverse family of early aquatic mammals of the order Cetacea. The family is named after paleocetologist Remington Kellogg.

<span class="mw-page-title-main">Jebel Qatrani Formation</span> Paleontological and geological formation in Faiyum Governorate, Egypt

The Jebel Qatrani Formation is a geologic formation located in the Faiyum Governorate of central Egypt. It is exposed between the Jebel Qatrani escarpment and the Qasr el Sagha escarpment, north of Birket Qarun lake near Faiyum. The formation conformably overlies the Qasr el Sagha Formation and is topped by the Widan el Faras Basalt. The age of the formation has been subject to debate, but the most recent research indicates that it covers both the latest parts of the Eocene and the Early Oligocene, spanning over the boundary between these two time periods.

The Qasr el Sagha Formation is a geological formation located in Egypt. The formation is part of the Wadi El Hitan World Heritage Site. The Qasr el Sagha Formation overlies the Birket Qarun Formation and is overlain by the Gebel Qatrani Formation. The sandstones and shales of the formation were deposited in a deltaic to shallow marine environment. It dates to the Late Eocene.

Protocetidae, the protocetids, form a diverse and heterogeneous group of extinct cetaceans known from Asia, Europe, Africa, South America, and North America.

<i>Saghacetus</i> Genus of mammals

Saghacetus is an extinct genus of basilosaurid early whale, fossils of which have been found in the Upper Eocene Qasr el Sagha Formation, Egypt.

Cordichelys is an extinct genus of podocnemidid turtle. It was around during the Eocene. Fossils of this turtle have been discovered at Wadi El Hitan as of November 2020.

Protosiren eothene is a species of Lutetian sirenian, found in the upper part of the Habib Rahi Formation, Pakistan.

Makaracetus is an extinct protocetid early whale the remains of which were found in 2004 in Lutetian layers of the Domanda Formation in the Sulaiman Range of Balochistan, Pakistan.

<i>Eosiren</i> Extinct genus of mammals

Eosiren is an extinct genus of sea cow that lived during the Late Eocene to Early Oligocene (Rupelian). Several fossils have been found in Egypt. It seems like the species E. abeli were contemporaneous with Protosiren and Eotheroides. like them, Eosiren closely resembled modern sirenians. It differs from them by having somewhat larger innominates and possess thigh bones.

Eocetus is an extinct protocetid early whale known from the early late Eocene Giushi Formation in Gebel Mokattam, outside Cairo, Egypt. The specimen was first named by Fraas as Mesocetus schweinfurthi. However, the name Mesocetus was previously used causing a change to the species name to Eocetus schweinfurthi. Since the genus was first described in the early 20th century, several other specimens, mostly isolated vertebrae, have been attributed to Eocetus, but the taxonomic status of these widely distributed specimens remain disputed.

<i>Prototherium</i>

Prototherium is a genus of extinct sirenian related to the dugong. It is known from middle (Bartonian) and upper Eocene deposits in Italy and Spain. Type species is P. veronenses

References

  1. 1 2 3 Gingerich, P. D.; Domning, D. P.; Blane, C. E.; Uhen, M. D. (1994). "Late Eocene sea cows (Mammalia, Sirenia) from Wadi Al Hitan in the Fayum Basin, Egypt". Contributions from the Museum of Paleontology. University of Michigan. 29 (2): 41–67.
  2. Samonds, K. E.; Zalmout, I. S.; Irwin, M. T.; Krause, D. W.; Rogers, R. R.; Raharivony, L. L. (2009). "Eotheroides lambondrano, new Middle Eocene seacow (Mammalia, Sirenia) from the Mahajanga Basin, Northwestern Madagascar". Journal of Vertebrate Paleontology. 29 (4): 1233–1243. Bibcode:2009JVPal..29.1233S. doi:10.1671/039.029.0417. S2CID   59466434.
  3. 1 2 "Ancient Pygmy Sea Cow Discovered". Science Daily. 18 December 2009. Retrieved 18 December 2009.
  4. Bajpai, S.; Thewissen, J. G. M.; Kapur, V. V.; Tiwari, B. N.; Sahni, A. (2006). "Eocene and Oligocene sirenians (Mammalia) from Kachchh, India". Journal of Vertebrate Paleontology. 26 (2): 400–410. doi:10.1671/0272-4634(2006)26[400:eaosmf]2.0.co;2. S2CID   86682899.
  5. Zdansky, O. (1138). "Eotherium majus sp. n., eine neue Sirene aus dem Mitteleozän von Ägypten". Palaeobiologica. 6: 429–434.
  6. 1 2 Iyad S. Zalmout and Philip D. Gingerich (2012). "Late Eocene Sea Cows (Mammalia, Sirenia) From Wadi Al Hitan In The Western Desert of Fayum, Egypt". University of Michigan Papers on Paleontology. 37: 1–158. hdl:2027.42/94568.
  7. Das, D. P.; Basu, P. K. (1994). "Study of Palaeogene marine mammals of Kachchh, Guarat". Geological Survey of India. 127 (2): 5.
  8. Gingerich, P. D. (1992). "Marine Mammals (Cetacean and Sirenia) from the Eocene of Gebel Mokattam and Fayum, Egypt: Stratigraphy, Age, and Paleoenvironments". Contributions from the Museum of Paleontology. University of Michigan. 30: 1–84.

Further reading