Libognathus

Last updated

Libognathus
Temporal range: Late Triassic
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Parareptilia
Order: Procolophonomorpha
Family: Procolophonidae
Tribe: Leptopleuronini
Genus: Libognathus
Small, 1997
Type species
Libognathus sheddi
Small, 1997

Libognathus (meaning "southwest jaw" in Greek) is an extinct genus of procolophonid parareptile from the Late Triassic of Texas. The type and only species, Libognathus sheddi, was named in 1997 from the Cooper Canyon Formation in the fossil-rich Post Quarry, which is found in Garza County. Libognathus was the first definite procolophonid discovered in the southwestern United States, although another possible procolophonid called Chinleogomphius was reported from the southwest before Libognathus was named. [1]

Description

Libognathus is known from the holotype specimen including the left dentary bone and coronoid process of the lower jaw, as well as six teeth implanted in the jaw. Recently, additional material including much of the anterior skull and several mandibles were referred to the genus. Libognathus can be identified as a procolophonid by its wide molar-like teeth, each of which has a ridge running along its width and several projections or cusps around it. It is considered a member of the procolophonid subfamily Leptopleuroninae based on the presence of three spine-like processes on the quadratojugal, the deep posterior and anterior margins of the dentary, the ventral deflection of the jaw articulation of the articular bone, and more conically shaped teeth at the front of the jaw. Libognathus is distinguished from other procolophonids by the steep angle of the lower margin of the dentary (indicating a deeper lower jaw than in related procolophonids such as Hypsognathus and Leptopleuron ) and a small hole or foramen on the jaw's outer surface below the fifth tooth. [1]

Several other jaw fragments have been identified as possible North American procolophonids, and have been given names such as Chinleogomphius, Xenodiphyodon , and Gomphiosauridion . The holotype Libognathus can be distinguished from these other remains by features in its teeth; while Chinleogomphius and Xenodiphyodon have three cusps per tooth, Libognathus has only two, and while Gomphiodon has a ridge positioned centrally on the tooth surface, the ridge on each tooth of Libognathus is positioned farther forward. [1]

Tooth replacement is evident on the inner surface of the jaw of Libognathus, where the exposed roots of teeth show smaller teeth pushing up from underneath them. This form of tooth replacement is similar to that in most other reptiles, but different from the condition in most other procolophonids, which have teeth that are fused to the bone of the jaw. [1]

Several procolophonid specimens from the Owl Rock Member of the Chinle Formation in Arizona have been found to be closely related, and possibly referrable to, Libognathus.

Related Research Articles

<i>Probainognathus</i> Extinct genus of cynodonts

Probainognathus meaning “progressive jaw” is an extinct genus of cynodonts that lived around 235 to 221.5 million years ago, during the Late Triassic in what is now Argentina. Together with the genus Bonacynodon from Brazil, Probainognathus forms the family Probainognathidae. Probainognathus was a relatively small, carnivorous or insectivorous cynodont. Like all cynodonts, it was a relative of mammals, and it possessed several mammal-like features. Like some other cynodonts, Probainognathus had a double jaw joint, which not only included the quadrate and articular bones like in more basal synapsids, but also the squamosal and surangular bones. A joint between the dentary and squamosal bones, as seen in modern mammals, was however absent in Probainognathus.

<i>Lesothosaurus</i> Extinct genus of ornithischian dinosaur

Lesothosaurus is a monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from Lesotho". The genus has only one valid species, Lesothosaurus diagnosticus. Lesothosaurus is one of the most completely-known early ornithischians, based on numerous skull and postcranial fossils from the Upper Elliot Formation. It had a simpler tooth and jaw anatomy than later ornithischians, and may have been omnivorous in some parts of the year.

<i>Prozostrodon</i> Extinct genus of cynodonts

Prozostrodon is an extinct genus of probainognathian cynodonts that was closely related to the ancestors of mammals. The remains were found in Brazil and are dated to the Carnian age of the Late Triassic. The holotype has an estimated skull length of 6.7 centimetres (2.6 in), indicating that the whole animal may have been the size of a cat. The teeth were typical of advanced cynodonts, and the animal was probably a carnivore hunting reptiles and other small prey.

<i>Omphalosaurus</i> Extinct genus of reptiles

Omphalosaurus is an extinct genus of marine reptile from the Early Triassic to Middle Triassic, thought to be in the order of Ichthyosauria. Most of what is known about Omphalosaurus is based on multiple jaw fragments, ribs, and vertebrae. Specimens of Omphalosaurus have been described from the western United States, Poland, Austria and the island of Spitsbergen off the northern coast of Norway.

<i>Gobiconodon</i> Extinct genus of mammals

Gobiconodon is an extinct genus of carnivorous mammals belonging to the family Gobiconodontidae. Undisputed records of Gobiconodon are restricted to the Early Cretaceous of Asia and North America, but isolated teeth attributed to the genus have also been described from formations in England and Morocco dating as far back as the Middle Jurassic. Species of Gobiconodon varied considerably in size, with G. ostromi, one of the larger species, being around the size of a modern Virginia opossum. Like other gobiconodontids, it possessed several speciations towards carnivory, such as shearing molariform teeth, large canine-like incisors and powerful jaw and forelimb musculature, indicating that it probably fed on vertebrate prey. Unusually among predatory mammals and other eutriconodonts, the lower canines were vestigial, with the first lower incisor pair having become massive and canine-like. Like the larger Repenomamus there might be some evidence of scavenging.

<i>Brasilodon</i> Extinct genus of mammaliamorphs

Brasilodon is an extinct genus of small, mammal-like cynodonts that lived in what is now Brazil during the Norian age of the Late Triassic epoch, about 225.42 million years ago. While no complete skeletons have been found, the length of Brasilodon has been estimated at 12 centimetres (4.7 in). Its dentition shows that it was most likely an insectivore. The genus is monotypic, containing only the species B. quadrangularis. Brasilodon belongs to the family Brasilodontidae, whose members were some of the closest relatives of mammals, the only cynodonts alive today. Two other brasilodontid genera, Brasilitherium and Minicynodon, are now considered to be junior synonyms of Brasilodon.

<i>Platyognathus</i> Extinct genus of reptiles

Platyognathus is an extinct genus of protosuchian crocodylomorph. Fossils are known from the Early Jurassic Lower Lufeng Formation in Yunnan, China and belong to the type and only species, P. hsui.

Variodens is an extinct genus of trilophosaur. Fossils have been found from the Emborough Quarries in the Mendip Hills of Somerset, England. These fossils have been uncovered from a Late Triassic fissure fill within Carboniferous-age limestone. The type and only known species is V. inopinatus, named in 1957.

<i>Leonerasaurus</i> Genus of sauropodomorph dinosaur from the Early Jurassic period

Leonerasaurus is a basal genus of sauropodomorph dinosaur. Currently, there is only one species known, named L. taquetrensis by Diego Pol, Alberto Garrido and Ignacio A. Cerda in 2011. The fossil, an incomplete subadult individual, was found in the Las Leoneras Formation in Argentina. This formation is probably Early Jurassic in age. Leonerasaurus was a small non-sauropod sauropodomorph, showing an unusual combination of basal and derived characters. This indicates that the evolution of early sauropodomorphs witnessed a great degree of convergent evolution.

Arctotraversodon is an extinct genus of traversodontid cynodonts from the Late Triassic of Canada. Fossils first described from the Wolfville Formation in Nova Scotia in 1984 represented the first known traversodontid from North America. The type and only species is A. plemmyridon and is represented by teeth and several dentary bones.

<i>Diodorus scytobrachion</i> Extinct species of reptile

Diodorus is a genus of silesaurid dinosauromorph that lived during the Late Triassic in what is now Morocco. Fossils were discovered in the Timezgadiouine Formation of the Argana Basin, and were used to name the new genus and species Diodorus scytobrachion. The genus name honors the mythological king Diodorus and the ancient historian Diodorus Siculus; the specific name is ancient Greek for 'leathery arm' and also honors the mythographer Dionysius Scytobrachion. The holotype specimen is a partial dentary bone (front of the lower jaw), and assigned specimens include isolated teeth, two humeri (upper arm bones), a metatarsal (foot bone), and femur (thigh bone).

<i>Cricodon</i> Extinct genus of cynodonts

Cricodon is an extinct genus of trirachodontid cynodonts that lived during the Early Triassic and Middle Triassic periods of Africa. A. W. Crompton named Cricodon based on the ring-like arrangement of the cuspules on the crown of a typical postcanine tooth. The epithet of the type species, C. metabolus, indicates the change in structure of certain postcanines resulting from replacement.

<i>Abdalodon</i> Extinct genus of cynodonts

Abdalodon is an extinct genus of late Permian cynodonts, known by its only species A. diastematicus.Abdalodon together with the genus Charassognathus, form the clade Charassognathidae. This clade represents the earliest known cynodonts, and is the first known radiation of Permian cynodonts.

Soumyasaurus is a small silesaurid dinosauriform from the Late Triassic (Norian) Cooper Canyon Formation of western Texas.

<i>Arisierpeton</i> Extinct genus of synapsids

Arisierpeton is an extinct genus of synapsids from the Early Permian Garber Formation of Richards Spur, Oklahoma. It contains a single species, Arisierpeton simplex.

<i>Kwanasaurus</i> Extinct genus of silesaurid dinosauromorph reptiles

Kwanasaurus is an extinct genus of silesaurid dinosauromorph reptiles from the Late Triassic of Colorado. It is known from a single species, Kwanasaurus williamparkeri. Kwanasaurus had a deeper, stronger skull and greater specialization for herbivory compared to other silesaurids. It also possessed many unique characteristics of the snout, ilium, and lower part of the femur. It was described along with new specimens of Dromomeron from the Eagle Basin, the northernmost extent of the Chinle Formation.

<i>Bonacynodon</i> Extinct genus of cynodonts

Bonacynodon is an extinct genus of cynodonts that lived in what is now southern Brazil during the Triassic period. The genus is monotypic, containing only the type species Bonacynodon schultzi. B. schultzi is known from two specimens, consisting of two partial skulls and some badly preserved parts of the postcranium. Both specimens were recovered from the Pinheiros-Chiniquá Sequence, part of the Santa Maria Supersequence of the Paraná Basin. This sequence preserves a faunal association known as the Dinodontosaurus Assemblage Zone, which contains numerous other species of cynodonts, dicynodonts and reptiles. Bonacynodon was a small, likely insectivorous cynodont, whose length has been estimated at around 30 centimetres (12 in). It can be distinguished from other cynodonts by its large, serrated (saw-like) canine teeth. Together with the genus Probainognathus of Argentina, it made up the family Probainognathidae, one of the earliest-diverging lineages of the clade Probainognathia. It was a fairly close relative of mammals, the only group of cynodonts alive today.

<i>Kalaallitkigun</i> Extinct genus of mammaliaforms

Kalaallitkigun is an extinct genus of haramiyidan mammaliaforms from the Late Triassic of Greenland. It contains a single species, Kalaallitkigun jenkinsi, which was described in 2020 from a partial dentary found in the Fleming Fjord Formation. More specifically, it was found in the mid-late Norian Carlsberg Fjord beds of the Ørsted Dal Member. It is the oldest of several mammaliaform species discovered in the Late Triassic sediments of Greenland. It is also the oldest mammaliaform with double-rooted teeth, and its pattern of tooth cusps help to clarify the evolution of haramiyidan teeth relative to their morganucodont-like ancestors.

<i>Kyhytysuka</i> Extinct marine reptile

Kyhytysuka is an extinct genus of ophthalmosaurian ichthyosaur from Early Cretaceous Colombia. The animal was previously assigned to the genus Platypterygius, but given its own genus in 2021. Kyhytysuka was a mid-sized ophthalmosaurian with heterodont dentition and several adaptations suggesting that it was a macropredatory vertebrate hunter living in shallow waters. It contains a single species, Kyhytysuka sachicarum.

<i>Santacruzgnathus</i> Extinct genus of cynodonts

Santacruzgnathus is an extinct genus of small cynodonts from the Late Triassic (Carnian) Santacruzodon Assemblage Zone of Brazil. It contains one species, S. abdalai. Santacruzgnathus is known from a single partial lower jaw with four postcanine teeth, only one of which is well-preserved. Some features of the specimen, including the slender shape of the jaw and the incipiently double-rooted teeth, indicate that the animal was an early member of Prozostrodontia, a group that includes mammals and their close relatives.

References

  1. 1 2 3 4 Small, B. J. (1997). "A new procolophonid from the Upper Triassic of Texas, with a description of tooth replacement and implantation". Journal of Vertebrate Paleontology. 17 (4): 674–678. doi:10.1080/02724634.1997.10011016.

2. Mueller, B. D., Small, B. J., Jenkins, X., Huttenlocker, A. K., & Chatterjee, S. (2023). Cranial anatomy of Libognathus sheddi Small, 1997 (Parareptilia, Procolophonidae) from the Upper Triassic Dockum Group of West Texas, USA. The Anatomical Record, 1–21. https://doi.org/10.1002/ar.25364