Theledectes Temporal range: Middle Triassic, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | † Parareptilia |
Order: | † Procolophonomorpha |
Family: | † Procolophonidae |
Subfamily: | † Theledectinae |
Genus: | † Theledectes Modesto & Damiani, 2003 |
Type species | |
†Theledectes perforatus (Gow, 1977a [originally Thelegnathus perforatus]) |
Theledectes is an extinct genus of theledectine procolophonid parareptile from middle Triassic (early Anisian stage) deposits of Free State Province, South Africa.The type species, Theledectes perforatus, is based on the holotype BP/1/4585, a flattened skull. This skull was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Rouxville District and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group (Karoo Basin). The genus was first named by Sean P. Modesto and Ross J. Damiani in 2003. [1] However, the species was initially assigned to the genus Thelegnathus (now considered to be a nomen dubium ) by C.E. Gow in 1977, as the species Thelegnathus perforatus. [2]
Most of the skull is poorly preserved, with the exception of the tooth-bearing elements. Theledectes is the only known parareptile with more than one row of marginal teeth (teeth along the edge of the mouth). The only other reptiles with multiple marginal tooth rows are rhynchosaurs and some captorhinids. The lower jaw has two large incisor-like teeth, akin to Scoloparia . However, the dentition is also distinctly inset from the rest of the face by a large curved area, a trait also known in Hypsognathus . [1] [2] The teeth are large and blunted cones, with circular bases, in contrast to most procolophonids which have teeth that are wider than long. Similar teeth are also observed in "Eumetabolodon" dongshengensis, a Chinese procolophonid previously considered a species of Eumetabolodon . In 2008, Juan Carlos Cisneros named the subfamily Theledectinae to encompass Theledectes and "E." dongshengensis. [3]
Cladogram after Cisneros, 2008: [3]
Procolophonidae |
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Milleretta is an extinct genus of millerettid parareptile from the Late Permian of South Africa. Fossils have been found in the Balfour Formation. Milleretta was a moderately sized, lizard-like animal, about 60 centimetres (24 in) in length. It was probably insectivorous. Its only known species is Milleretta rubidgei, making Milleretta a monospecific genus.
Procolophon is a genus of lizard-like procolophonid parareptiles that first appeared in the Early Triassic (Induan) of South Africa, Brazil, and Antarctica. It persisted through the Permian–Triassic extinction event, but went extinct in the beginning of the Early Middle Triassic. The type species is P. trigoniceps.
Leptopleuron is an extinct genus of procolophonid that lived in the dry lands during the late Triassic in Elgin of northern Scotland and was the first to be included in the clade of Procolophonidae. First described by English paleontologist and biologist Sir Richard Owen, Leptopleuron is derived from two Greek bases, leptos for "slender" and pleuron for "rib," describing it as having slender ribs. The fossil is also known by a second name, Telerpeton, which is derived from the Greek bases tele for "far off" and herpeton for "reptile." In Scotland, Leptopleuron was found specifically in the Lossiemouth Sandstone Formation. The yellow sandstone it was located in was poorly lithified with wind coming from the southwest. The environment is also described to consist of barchan dunes due to the winds, ranging up to 20 m tall that spread during dry phases into flood plains. Procolophonoids such as Leptopleuron were considered an essential addition to the terrestrial ecosystem during the Triassic.
Microposaurus is an extinct genus of trematosaurid temnospondyl. Fossils are known from the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Rouse Hill Siltstone of Australia that date back to the Anisian stage of the Middle Triassic. These aquatic creatures were the short snouted lineage from Trematosaurinae.
Acleistorhinus (ah-kles-toe-RYE-nuss) is an extinct genus of parareptile known from the Early Permian of Oklahoma. It is notable for being the earliest known anapsid reptile yet discovered. The morphology of the lower temporal fenestra of the skull of Acleistorhinus bears a superficial resemblance to that seen in early synapsids, a result of convergent evolution. Only a single species, A. pteroticus, is known, and it is classified in the Family Acleistorhinidae, along with Colobomycter.
Owenetta is an extinct genus of owenettid procolophonian parareptile. Fossils have been found from the Beaufort Group in the Karoo Basin of South Africa. Although most procolophonians lived during the Triassic, Owenetta existed during the Wuchiapingian and Changhsingian stages of the Late Permian as well as the early Induan stage of the Early Triassic. It is the type genus of the family Owenettidae, and can be distinguished from other related taxa in that the posterior portion of the supratemporal bears a lateral notch and that the pineal foramen is surrounded by a depressed parietal surface on the skull table.
Procolophonoidea is an extinct superfamily of procolophonian parareptiles. Members were characteristically small, stocky, and lizard-like in appearance. Fossils have been found worldwide from many continents including Antarctica. The first members appeared during the Late Permian in the Karoo Basin of South Africa.
Phonodus is an extinct genus of procolophonid parareptile. It is known from a single skull found from the Early Triassic Katberg Formation in South Africa. It is the oldest known member of the subfamily Leptopleuroninae, and was likely the result of a procolophonid migration into the Karoo Basin from Laurasia after the Permo-Triassic extinction event. Because Phonodus had large maxillary teeth underneath a large antorbital buttress, and a lack of ventral temporal emargination along the side of the skull, it probably had a durophagous diet.
Kitchingnathus is an extinct genus of basal procolophonid parareptile from Early Triassic deposits of Eastern Cape Province, South Africa. It is known from the holotype BP/1/1187, skull and partial postcranium, which was first assigned to the more derived Procolophon trigoniceps. It was collected by the South African palaeontologist, James W. Kitching in October 1952 from Hobbs Hill, west of Cathcart. It was found in the middle or upper part of the Katberg Formation of the Beaufort Group and referred to the uppermost Lystrosaurus Assemblage Zone. It was first named by Juan Carlos Cisneros in 2008 and the type species is Kitchingnathus untabeni. The generic name honours James W. Kitching, and "gnathus", from Greek gnathos meaning mandible. The specific name meaning "from the hill", in isiZulu, is in reference to the locality where the fossil was found.
Sauropareion is an extinct genus of basal procolophonid parareptile from earliest Triassic deposits of Eastern Cape Province, South Africa. It is known from the holotype SAM PK-11192, skull and partial postcranium. It was collected by the late L. D. Boonstra in 1935 from Barendskraal in the Middelburg District and referred to the Lystrosaurus Assemblage Zone of the Beaufort Group. It was first named by Sean P. Modesto, Hans-Dieter Sues and Ross J. Damiani in 2001 and the type species is Sauropareion anoplus. The generic name means "lizard", sauros, and "cheek", pareion from Greek in reference to the lizard-like appearance of the temporal region. The specific name comes from the Greek word anoplos, meaning "without arms or armour".
Coletta is an extinct genus of basal procolophonid parareptile from Early Triassic deposits of Eastern Cape Province, South Africa. It is known from the holotype GHG 228, a skull with fragmentary lower jaws. It was collected on the farm Brakfontein 333 in the Cradock District. It was found in the Katberg Formation of the Beaufort Group and referred to the Lystrosaurus Assemblage Zone. It was first named by Christopher E. Gow in 2000 and the type species is Coletta seca.
Pintosaurus is an extinct genus of basal procolophonid parareptile from Late Triassic deposits of northeastern Uruguay. It is known from the holotype FC-DPV 1181, a partial skull. It was collected from the Buena Vista Formation of the Paraná Basin, in Colonia Orozco, Cerro Largo Department. It was first named by Graciela Piñeiro, Alejandra Rojas and Martín Ubilla in 2004 and the type species is Pintosaurus magnidentis. The generic name honours Dr. Iraja Damiani Pinto. The specific name means "with a large tooth" in Latin, a reference to the large palatal tooth pair.
Theledectinae is an extinct subfamily of parareptiles within the family Procolophonidae. Theledectines existed in South Africa, China and Australia during the Early-Middle Triassic period. Theledectinae was named by Juan Carlos Cisneros in 2008 to include the genus Theledectes, and the species "Eumetabolodon" dongshengensis. "E." dongshengensis represents a new genus from China. Cladistically, it is defined as "All taxa more closely related to Theledectes perforatus than to Procolophon trigoniceps Owen, 1876". In 2020, Hamley add the new genus Eomurruna from Australia to this subfamily
Phaanthosaurus is an extinct genus of basal procolophonid parareptile from early Triassic deposits of Nizhnii Novgorod, Russian Federation. It is known from the holotype PIN 1025/1, a mandible. It was collected from Vetluga River, Spasskoe village and referred to the Vokhmian terrestrial horizon of the Vokhma Formation. It was first named by P. K. Chudinov and B. P. Vjushkov in 1956 and the type species is Phaanthosaurus ignatjevi.
Teratophon is an extinct genus of procolophonine procolophonid parareptile from middle Triassic deposits of Free State Province, South Africa. It is known from the holotype BP/1/4299, a nearly complete skull. It was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Rouxville District and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group. It was first named by Sean P. Modesto and Ross J. Damiani in 2003 and the type species is Teratophon spinigenis. It was first assigned to a species of Thelegnathus, Thelegnathus spinigenis.
Thelephon is an extinct genus of procolophonine procolophonid parareptile from middle Triassic deposits of Free State Province, South Africa. It is known from the holotype BP/1/3512, a partial skull lacking the snout and anterior third of the mandible. It was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Winnaarsbaken and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group. It was first named by Sean P. Modesto and Ross J. Damiani in 2003 and the type species is Thelephon contritus. It was first assigned to a species of Thelegnathus, Thelegnathus contritus.
Thelerpeton is an extinct genus of procolophonine procolophonid parareptile from middle Triassic deposits of Free State Province, South Africa. It is known from the holotype BP/1/4538, a nearly complete skull. It was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Rouxville District and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group. It was first named by Sean P. Modesto and Ross J. Damiani in 2003 and the type species is Thelerpeton oppressus. It was first assigned to a species of Thelegnathus, Thelegnathus oppressus.
Eumetabolodon is an extinct genus of procolophonine procolophonid parareptile from early and middle Triassic deposits of Nei Mongol, northern China. Two species of Eumetabolodon were named by J. L. Li in 1983 and the type species is Eumetabolodon bathycephalus.
Palacrodon is an extinct genus of Triassic reptile with a widespread distribution. It was initially described from teeth collected in Early Triassic deposits in South Africa, and later reported from the Early Triassic of Antarctica and the Late Triassic of Arizona. Although previously considered an early rhynchocephalian, it is currently considered to be a non-saurian neodiapsid.
Eomurruna is a genus of procolophonid reptile that existed in what is now Queensland, Australia during the Early Triassic period. The genus is made up of a single species, E. yurrgensis, originally uncovered within the Arcadia Formation in 1985. Since then over 40 specimens have been referred to the genus, making Eomurruna one of the most complete organisms so far found from the Mesozoic of Australia.