Thelerpeton Temporal range: Middle Triassic, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | † Parareptilia |
Order: | † Procolophonomorpha |
Family: | † Procolophonidae |
Subfamily: | † Procolophoninae |
Genus: | † Thelerpeton Modesto & Damiani, 2003 |
Type species | |
†Thelerpeton oppressus (Gow, 1977 [originally Thelegnathus oppressus]) |
Thelerpeton is an extinct genus of procolophonine procolophonid parareptile from middle Triassic (early Anisian stage) deposits of Free State Province, South Africa. It is known from the holotype BP/1/4538, a nearly complete skull. It was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Rouxville District and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group (Karoo Basin). It was first named by Sean P. Modesto and Ross J. Damiani in 2003 and the type species is Thelerpeton oppressus. It was first assigned to a species of Thelegnathus (now considered to be a nomen dubium ), Thelegnathus oppressus. [1] [2]
In their phylogenetic analyses, Butler et al. (2023) defined Thelerpeton as a sister taxon to Teratophon and Procolophon or as a sister taxon to clade consisting of Teratophon and Procolophon within Procolophoninae. The results are shown in two cladograms below: [3]
Analyses 1 and 3: Strict consensus of 760 and 18 most parsimonious trees (MPTs).
| Analysis 2: Single MPT.
|
The Lystrosaurus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the upper Adelaide and lower Tarkastad Subgroups of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. This biozone has outcrops in the south central Eastern Cape and in the southern and northeastern Free State. The Lystrosaurus Assemblage Zone is one of eight biozones found in the Beaufort Group, and is considered to be Early Triassic in age.
The Cynognathus Assemblage Zone is a tetrapod biozone utilized in the Karoo Basin of South Africa. It is equivalent to the Burgersdorp Formation, the youngest lithostratigraphic formation in the Beaufort Group, which is part of the fossiliferous and geologically important Karoo Supergroup. The Cynognathus Assemblage Zone is the youngest of the eight biozones found in the Beaufort Group, and is considered to be late Early Triassic (Olenekian) to early Middle Triassic (Anisian) in age. The name of the biozone refers to Cynognathus crateronotus, a large and carnivorous cynodont therapsid which occurs throughout the entire biozone.
The Pristerognathus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the upper Abrahamskraal Formation and lowermost Teekloof Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching not more than 300 metres (980 ft), occur just east of Sutherland through to Beaufort West in the south and Victoria West in the north. Exposures are also found west of Colesberg and south of Graaff-Reinet. The Pristerognathus Assemblage Zone is the third biozone of the Beaufort Group.
Procolophon is a genus of lizard-like procolophonid parareptiles that first appeared in the Early Triassic (Induan) of South Africa, Brazil, and Antarctica. It persisted through the Permian–Triassic extinction event, but went extinct in the beginning of the Early Middle Triassic. The type species is P. trigoniceps.
Parotosuchus is an extinct genus of capitosaurian temnospondyl amphibians within the family Mastodonsauridae. Fossils are known from the Early Triassic of Europe, Africa, Australia, and Antarctica. It was about 2 metres (6.6 ft) long and likely lived in aquatic environments such as lakes and rivers. Parotosuchus was covered in a scaly skin, unlike the smooth skin of modern-day amphibians, and probably moved with an eel-like motion in the water.
Microposaurus is an extinct genus of trematosaurid temnospondyl. Fossils are known from the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Rouse Hill Siltstone of Australia that date back to the Anisian stage of the Middle Triassic. These aquatic creatures were the short snouted lineage from Trematosaurinae.
Vanastega is an extinct genus of Triassic temnospondyl amphibian in the family Brachyopidae. It is known from the Cynognathus Assemblage Zone in Burgersdorp, South Africa. The genus contains just one species, Vanastega plurimidens, the type species.
Owenetta is an extinct genus of owenettid procolophonian parareptile. Fossils have been found from the Beaufort Group in the Karoo Basin of South Africa. Although most procolophonians lived during the Triassic, Owenetta existed during the Wuchiapingian and Changhsingian stages of the Late Permian as well as the early Induan stage of the Early Triassic. It is the type genus of the family Owenettidae, and can be distinguished from other related taxa in that the posterior portion of the supratemporal bears a lateral notch and that the pineal foramen is surrounded by a depressed parietal surface on the skull table.
Owenettidae is an extinct family of procolophonian parareptiles. Fossils have been found primarily from Africa and Madagascar, with one genus present from South America. It is the sister taxon to the family Procolophonidae. Modesto and Damiani (2007) defined Owenettidae as a stem-based group including Owenetta rubidgei and all species closely related to it than to Procolophon trigoniceps.
Procolophonoidea is an extinct superfamily of procolophonian parareptiles. Members were characteristically small, stocky, and lizard-like in appearance. Fossils have been found worldwide from many continents including Antarctica. The first members appeared during the Late Permian in the Karoo Basin of South Africa.
Sauropareion is an extinct genus of basal procolophonid parareptile from earliest Triassic deposits of Eastern Cape Province, South Africa. It is known from the holotype SAM PK-11192, skull and partial postcranium. It was collected by the late L. D. Boonstra in 1935 from Barendskraal in the Middelburg District and referred to the Lystrosaurus Assemblage Zone of the Beaufort Group. It was first named by Sean P. Modesto, Hans-Dieter Sues and Ross J. Damiani in 2001 and the type species is Sauropareion anoplus. The generic name means "lizard", sauros, and "cheek", pareion from Greek in reference to the lizard-like appearance of the temporal region. The specific name comes from the Greek word anoplos, meaning "without arms or armour".
Coletta is an extinct genus of basal procolophonid parareptile from Early Triassic deposits of Eastern Cape Province, South Africa. It is known from the holotype GHG 228, a skull with fragmentary lower jaws. It was collected on the farm Brakfontein 333 in the Cradock District. It was found in the Katberg Formation of the Beaufort Group and referred to the Lystrosaurus Assemblage Zone. It was first named by Christopher E. Gow in 2000 and the type species is Coletta seca.
Theledectes is an extinct genus of theledectine procolophonid parareptile from middle Triassic deposits of Free State Province, South Africa.The type species, Theledectes perforatus, is based on the holotype BP/1/4585, a flattened skull. This skull was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Rouxville District and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group. The genus was first named by Sean P. Modesto and Ross J. Damiani in 2003. However, the species was initially assigned to the genus Thelegnathus by C.E. Gow in 1977, as the species Thelegnathus perforatus.
Theledectinae is an extinct subfamily of parareptiles within the family Procolophonidae. Theledectines existed in South Africa, China and Australia during the Early-Middle Triassic period. Theledectinae was named by Juan Carlos Cisneros in 2008 to include the genus Theledectes, and the species "Eumetabolodon" dongshengensis. "E." dongshengensis represents a new genus from China. Cladistically, it is defined as "All taxa more closely related to Theledectes perforatus than to Procolophon trigoniceps Owen, 1876". In 2020, Hamley add the new genus Eomurruna from Australia to this subfamily.
Teratophon is an extinct genus of procolophonine procolophonid parareptile from middle Triassic deposits of Free State Province, South Africa. It is known from the holotype BP/1/4299, a nearly complete skull. It was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Rouxville District and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group. It was first named by Sean P. Modesto and Ross J. Damiani in 2003 and the type species is Teratophon spinigenis. It was first assigned to a species of Thelegnathus, Thelegnathus spinigenis.
Thelephon is an extinct genus of procolophonine procolophonid parareptile from middle Triassic deposits of Free State Province, South Africa. It is known from the holotype BP/1/3512, a partial skull lacking the snout and anterior third of the mandible. It was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Winnaarsbaken and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group. It was first named by Sean P. Modesto and Ross J. Damiani in 2003 and the type species is Thelephon contritus. It was first assigned to a species of Thelegnathus, Thelegnathus contritus.
Procolophoninae is an extinct subfamily of procolophonid parareptiles from the late Early Triassic to the early Middle Triassic of Africa, Antarctica, Asia, Europe and South America. Currently, the oldest-known procolophonine is Procolophon from the earliest Olenekian stage.
Leptopleuroninae is an extinct subfamily of procolophonid reptiles. The oldest member of Leptopleuroninae is Phonodus dutoitorum from the Induan age of the Early Triassic. It is the only procolophonid group that survived into the Late Triassic.
Barasaurus is an extinct genus of owenettid procolophonoid parareptile known from the Late Permian and Early Triassic of Madagascar. It contains a single species, Barasaurus besairiei.
Saurodektes is an extinct genus of owenettid procolophonoid parareptile known from the earliest Triassic deposits of Eastern Cape Province, South Africa. It was first named by Sean P. Modesto, Ross J. Damiani, Johann Neveling and Adam M. Yates in 2003 and the type species is Saurodectes rogersorum. The generic name Saurodectes was preoccupied by the generic name of Saurodectes vrsanskyi Rasnitsyn & Zherikhin, 2000, a fossil chewing lice known from the Early Cretaceous of Russia. Thus, an alternative generic name, Saurodektes, was proposed by Modesto et al. in 2004. The generic name means "lizard", sauros, and "biter", dektes from Greek. The specific name, rogersorum, honors Richard and Jenny Rogers, owners of the farm Barendskraal, for their hospitality, support and interest in the work of the paleontologists who recovered the holotype. Saurodektes is known solely from the holotype BP/1/6025, a partial skull and some fragmentary partial postcranial elements, housed at the Bernard Price Institute for Palaeontological Research, although the unprepared specimens BP/1/6044, BP/1/6045 and BP/1/6047 might also be referable to it. All specimens were collected on the slopes of the Manhaar Hill at Barendskraal in the Middelburg District, from the Palingkloof Member of the Balfour Formation, Beaufort Group, only 12 metres below the base of the Katberg Formation. This horizon belongs to the Lystrosaurus Assemblage Zone, dating to the early Induan stage of the Early Triassic period.