Anthodon Temporal range: Changhsingian, | |
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Drawing of BMNH 47337 by Griesbach, 1876 | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | † Parareptilia |
Order: | † Procolophonomorpha |
Clade: | † Pareiasauria |
Clade: | † Pumiliopareiasauria |
Genus: | † Anthodon Owen, 1876 [1] |
Type species | |
†Anthodon serrarius | |
Other species | |
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Synonyms | |
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Anthodon (meaning "flower tooth") is an extinct genus of pareiasaur parareptile from the Permian period of South Africa and Tanzania.
In 1845, amateur geologists William Guybon Atherstone and Andrew Geddes Bain discovered several fossils near Dassieklip, Cape Province, in the Bushman's River Valley. [2] [3] This was the first dinosaur find in Africa and in the Southern Hemisphere. [4] In 1849 and 1853, Bain sent some of the fossils to palaeontologist Richard Owen for identification. Among them was an upper jaw Bain referred to as the "Cape Iguanodon ", so the site was named "Iguanodonhoek". Atherstone published a short paper about the discovery in 1857, [2] but lamented in 1871 that it had thus far received no attention in London. [3] [5] In 1876 Owen named a series of specimens from the collection Anthodon serrarius, basing the generic name on the resemblance of the teeth to a flower. [3] [6] The partial holotype skull BMNH 47337, the left jaw BMNH 47338, the matrix BMNH 47338 including bone fragments and impressions of the anterior skull, and the vertebrae BMNH 47337a were all assigned to Anthodon. [7] In 1882, Othniel Charles Marsh assigned Anthodon to Stegosauridae based on BMNH 47338, and in 1890, Richard Lydekker found that although Anthodon was a pareiasaur, its teeth were similar to those of the Stegosauridae. [7] Lydekker in 1890 also corrected a mistake of Owen, who had incorrectly summarised all the material as coming from a single locality, whereas there was separate material from two clearly distinct localities. [3]
Richard Owen, who described Anthodon, thought it was a dinosaur because dinosaurian skull material from the Early Cretaceous had become associated with the Permian material. The dinosaur material was later separated out by Robert Broom in 1912 and was renamed as the stegosaurid Paranthodon by Franz Nopcsa in 1929. [8]
A possible second species, A. minusculus, was named by Sidney Haughton in 1932 based on remains found in the Cistecephalus other zone of the Usili Formation in Tanzania. [9] Later authors have suggested that A. minusculus may have been the same animal as A. serratus. [10]
Pareiasaurus parvus (Haughton, 1913) and Propappus parvus (Haughton, 1913) were also synonymised with Anthodon serrarius. [10] The holotype of both species was SAM 2351, a pelvis discovered near Dunedin, Western Cape, South Africa. [11]
Anthodon combines the primitive feature of interpterygoid fenestrae with an advanced feature of turtle-like armor. It was about 1.2 to 1.5 meters (3.9 to 4.9 ft) in length, and weighed around 80 to 100 kg (180 to 220 lb). Small dermal ossicles covered the body, while the pattern of armor plates on the back reminiscent of a turtle shell. The tail was further shortened relative to less derived forms. Some other forms are characterized by having smooth skulls and armor on the dorsal midline. [6]
The skull was small, and the cheekbones unornamented as in other pareiasaurs. [12] The skull is 30 cm in length and quite lightly built. The cheekbones form very large quadratojugal "horns" that extend downwards to a great degree, but with a smooth unornamented surface. The mandible has ventral protrusions (further "horns"). The postparietals are fused and, along with the tabulars, located on the skull roof, as in more primitive diadectomorphs. There are 11 to 14 pairs of overlapping teeth, of small and uniform size, each with 8 to 15 cusps, giving them, as with all pareiasaurs, a leaf-like or flower like appearance, hence the generic name "flower tooth". [6]
Andrew Geddes Bain, was a South African geologist, road engineer, palaeontologist and explorer.
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivores that typically bore a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers.
Bradysaurus was a large, early and common pareiasaur. They possessed a covering of armoured scutes, likely serving as defense against their main predators, the gorgonopsians.
Pelorosaurus is a genus of titanosauriform sauropod dinosaur. Remains referred to Pelorosaurus date from the Early Cretaceous period, about 140-125 million years ago, and have been found in England and Portugal. Thomas Holtz estimated its length at 24 meters.
Moschops is an extinct genus of therapsids that lived in the Guadalupian epoch, around 265–260 million years ago. They were heavily built plant eaters, and they may have lived partly in water, as hippopotamuses do. They had short, thick heads and might have competed by head-butting each other. Their elbow joints allowed them to walk with a more mammal-like gait rather than crawling. Their remains were found in the Karoo region of South Africa, belonging to the Tapinocephalus Assemblage Zone. Therapsids, such as Moschops, are synapsids, the dominant land animals in the Permian period, which ended 252 million years ago.
Massospondylus was a genus of sauropodomorph dinosaur from the Early Jurassic. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been named. Fossils have since been found at other locations in South Africa, Lesotho, and Zimbabwe. Material from Arizona's Kayenta Formation, India, and Argentina has been assigned to the genus at various times, but the Arizonan and Argentinian material are now assigned to other genera.
Gorgonopsia is an extinct clade of sabre-toothed therapsids from the Middle to Upper Permian roughly 265 to 252 million years ago. They are characterised by a long and narrow skull, as well as elongated upper and sometimes lower canine teeth and incisors which were likely used as slashing and stabbing weapons. Postcanine teeth are generally reduced or absent. For hunting large prey, they possibly used a bite-and-retreat tactic, ambushing and taking a debilitating bite out of the target, and following it at a safe distance before its injuries exhausted it, whereupon the gorgonopsian would grapple the animal and deliver a killing bite. They would have had an exorbitant gape, possibly in excess of 90°, without having to unhinge the jaw.
Cistecephalus is an extinct genus of dicynodont therapsid from the Late Permian of southern Africa. It was a small, specialised, burrowing dicynodont, possibly with habits similar to a modern mole. The head was flattened and wedge-shaped, the body long, and the forelimbs very strong, with similarities in structure to the forelimb of modern burrowing mammals.
Algoasaurus is a genus of sauropod dinosaur from the Berriasian-early Valanginian-age Early Cretaceous Upper Kirkwood Formation of Cape Province, South Africa, specifically near a town called Despatch. Only one species, A. bauri, is known.
Paranthodon is a genus of stegosaurian dinosaur that lived in what is now South Africa during the Early Cretaceous, between 139 and 131 million years ago. Discovered in 1845, it was one of the first stegosaurians found. Its only remains, a partial skull, isolated teeth, and fragments of vertebrae, were found in the Kirkwood Formation. British paleontologist Richard Owen initially identified the fragments as those of the pareiasaur Anthodon. After remaining untouched for years in the British Museum of Natural History, the partial skull was identified by South African paleontologist Robert Broom as belonging to a different genus; he named the specimen Palaeoscincus africanus. Several years later, Hungarian paleontologist Franz Nopcsa, unaware of Broom's new name, similarly concluded that it represented a new taxon, and named it Paranthodon owenii. Since Nopcsa's species name was assigned after Broom's, and Broom did not assign a new genus, both names are now synonyms of the current binomial, Paranthodon africanus. The genus name combines the Ancient Greek para (near) with the genus name Anthodon, to represent the initial referral of the remains.
Regnosaurus is a genus of herbivorous stegosaurian dinosaur that lived during the Early Cretaceous Period in what is now England. It was one of the first stegosaurs discovered.
Ornithopsis is a genus of sauropod dinosaur, from the Early Cretaceous of England. The type species, which is the only species seen as valid today, is O. hulkei, which is only known from fragmentary remains, and has been regarded by many authors as dubious.
Gorgonops is an extinct genus of gorgonopsian therapsid, of which it is the type genus. Gorgonops lived during the Late Permian (Wuchiapingian), about 260–254 million years ago in what is now South Africa.
Palaeoscincus is a dubious genus of ankylosaurian dinosaur based on teeth from the mid-late Campanian-age Upper Cretaceous Judith River Formation of Montana. Like several other dinosaur genera named by Joseph Leidy, it is an historically important genus with a convoluted taxonomy that has been all but abandoned by modern dinosaur paleontologists. Because of its wide use in the early 20th century, it was somewhat well known to the general public, often through illustrations of an animal with the armor of Edmontonia and the tail club of an ankylosaurid.
Lycorhinus is a genus of heterodontosaurid ornithischian dinosaur from the Early Jurassic strata of the Elliot Formation located in the Cape Province, South Africa.
Rhinesuchus is a large temnospondyl. Remains of the genus are known from the Permian of the South African Karoo Basin's Tapinocephalus and Cistecephalus assemblage zones, both belonging to the Beaufort Group. The skull of Rhinesuchus had a flat triangular shape with blunt snout similar to some of the other large temnospondyls, and had a palate filled with small sharp teeth, suggesting that it hunted fish. Also, the small eyes were on top of the head suggesting that it approached its prey from below.
Suchosaurus is a spinosaurid dinosaur from Cretaceous England and Portugal, originally believed to be a genus of crocodile. The type material, consisting of teeth, was used by British palaeontologist Richard Owen to name the species S. cultridens in 1841. Later in 1897, French palaeontologist Henri-Émile Sauvage named a second species, S. girardi, based on two fragments from the mandible and one tooth discovered in Portugal. Suchosaurus is possibly a senior synonym of the contemporary spinosaurid Baryonyx, but is usually considered a dubious name due to the paucity of its remains, and is considered an indeterminate baryonychine. In the Wadhurst Clay Formation of what is now southern England, Suchosaurus lived alongside other dinosaurs, as well as plesiosaurs, mammals, and crocodyliforms.
Lydekkerina is an extinct genus of stereospondyl temnospondyl. It is the type genus of the family Lydekkerinidae. Fossils have been collected from Early Triassic deposits in South Africa and Australia. The type species is L. huxleyi, first described in 1889. While most other stereospondyls were semiaquatic, Lydekkerina was exclusively terrestrial.
Scymnosaurus is a dubious genus of therocephalian therapsids based upon various fossils of large early therocephalians. The genus was described by Robert Broom in 1903 with S. ferox, followed by S. watsoni in 1915 and a third, S. major, by Lieuwe Dirk Boonstra in 1954. Each of these species are considered nomen dubia today and based upon specimens belonging to two separate families of therocephalians. S. ferox and S. major represent specimens of Lycosuchidae incertae sedis, while S. watsoni is Scylacosauridae incertae sedis. Broom named a fourth species in 1907 from KwaZulu-Natal, S. warreni, though he later referred it to Moschorhinus as a valid species in 1932 but now is recognised as being synonymous with M. kitchingi.
Embrithosaurus was a pareiasaur from the Permian of South Africa.