List of Odonata species of Sri Lanka

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Sri Lanka is a tropical island situated close to the southern tip of India. The invertebrate fauna is as large as it is common to other regions of the world. There are approximately 2 million known species of arthropods, and this number continues to grow. Thus, it is difficult to determine the exact number of Odonata species within particular regions. The following is a list of the dragonflies and damselflies of Sri Lanka.

Dragonflies & Damselflies

Phylum: Arthropoda
Class: Insecta
Order: Odonata.

Dragonflies are predators. The larvae inhabit water and adults fly near aquatic places. They are diverse in color and shape. There are major 2 types of odonates in the world; they are dragonflies and damselflies. The eyes are closer together in dragonflies, and their wings are held broadly opened from the body. They are robust in nature. In contrast, damselflies are delicately built small odonates, with well separated compound eyes. During rest, they do not expand their wings; the wings are folded over the abdomen or slightly spread.

Damselflies are categorized in to Suborder: Zygoptera; and dragonflies into Suborder: Anisoptera. 131 described species within 13 families can be found in Sri Lanka, with three new species in 2016. [1] [2]

The 65 endemic species and marked with an asterisk (*).

Suborder: Zygoptera - Damselflies

Damselflies are insect s of suborder Zygoptera in the order Odonata. They are similar to dragonflies, which constitute the other odonatan suborder, Anisoptera, but are smaller, have slimmer bodies, and most species fold the wings along the body when at rest. An ancient group, damselflies have existed since at least the Lower Permian, and are found on every continent except Antarctica.

All damselflies are predatory; both nymphs and adults eat other insects. The nymphs are aquatic, with different species living in a variety of freshwater habitats including acid bogs, ponds, lakes and rivers. The nymphs moult repeatedly, at the last moult climbing out of the water to undergo metamorphosis. The skin splits down the back, they emerge and inflate their wings and abdomen to gain their adult form. Their presence on a body of water indicates that it is relatively unpolluted, but their dependence on freshwater makes them vulnerable to damage to their wetland habitats.

64 Damselfly species can be found in Sri Lanka.

Family: Calopterygidae - Jewelwings

Common nameBinomialLocal subspeciesMaleFemale
Stream glory Neurobasis chinensisssp.chinensis Neurobasis chinensis-Thattekad-2015-09-13-001.jpg Neurobasis chinensis-Aralam-2016-10-29-003.jpg
Black-tipped forest glory Vestalis apicalisssp.nigrescens* Black Tipped Forest Glory (Vestalis apicalis) male. (44158153825).jpg Black Tipped Forest Glory (Vestalis apicalis)female., cutttticcirrkn tnnlttumpi. (40686293970).jpg

Family: Chlorocyphidae - Jewels

Common nameBinomialLocal subspeciesMaleFemale
Adam's gem Libellago adami*
Ultima gem Libellago finalis*
Green's gem Libellago greeni* Libellago greeni male.JPG Libellago greeni female.JPG
Corbett's gem Libellago corbeti*

Family: Euphaeidae - Gossamerwings

Common nameBinomialLocal subspeciesMaleFemale
Shining gossamerwing Euphaea splendens*

Family: Lestidae - Spreadwings

Common nameBinomialLocal subspeciesMaleFemale
Sri Lanka reedling Indolestes divisus*
Mountain reedling Indolestes gracilisssp.gracilis* Davidraju IMG 9734.jpg Indolestes gracilis - female.jpg
Emerald spreadwing Lestes elatus Lestes elatus-Kadavoor-2016-07-04-004.jpg Lestes elatus-Madayippara.jpg
Malabar spreadwing Lestes malabaricus Lestes malabaricus Fraser (cropped).jpg
Scalloped spreadwing Lestes praemorsusssp.decipiens Lestes praemorsus male.jpg
Emerald Sri Lanka spreadwing Sinhalestes orientalis*

Family: Coenagrionidae - Narrow-winged damselflies

Common nameBinomialLocal subspeciesMaleFemale
Green striped slender dartlet Aciagrion occidentale Aciagrion occidentale 2 at Kadavoor.jpg Aciagrion occidentale-Kadavoor-2017-05-08-002.jpg
Variable wisp Agriocnemis feminassp.femina Variable wisp (Agriocnemis femina).jpg
Pygmy wisp Agriocnemis pygmaea Agriocnemis pygmaea male, Burdwan, West Bengal, India 14 03 2012.jpg Agriocnemis pygmaea female, Burdwan, West Bengal, India 11 10 2012.jpg
Azure dartlet Amphiallagma parvum Amphiallagma parvum by Davidraju IMG 9994.jpg Azure Dartlet ( Amphiallagma parvum) Female.jpg
Lieftinck's sprite Archibasis lieftincki* Archibasis lieftincki.jpg
Long-banded bluetail Archibasis oscillansssp. hanwellanensis* Archibasis oscillans-Kadavoor-2016-07-03-001.jpg Archibasis oscillans female (22617529972).jpg
Orange-tailed marsh dart Ceriagrion cerinorubellum Orange-tailed marsh dart 08030.jpg Davidraju IMG 7768.jpg
Coromandel marsh dart Ceriagrion coromandelianum Ceriagrion coromandelianum male, Burdwan, West Bengal, India 09 07 2013.jpg Ceriagrion coromandelianum, Burdwan, West Bengal, India 23 09 2012 (1).JPG
Golden dartlet Ischnura aurorassp.aurora Ischnura aurora01.jpg
Marsh bluetail Ischnura senegalensis Ischnura senegalensis(Male , Japan , 2018.09.02).jpg Ischnura senegalensis(Female,Japan,2018.10.08).jpg


Sri Lanka midget Mortonagrion ceylonicum*
Three striped blue dart Pseudagrion decorum Pseudagrion decorum-Thekkady-2016-12-02-001.jpg Pseudagrion decorum female by Manoj V Nair.jpg
Malabar sprite Pseudagrion malabaricum Pseudagrion malabaricum male.jpg Pseudagrion malabaricum Female.jpg
Blue riverdamsel Pseudagrion microcephalum Pseudagrion microcephalum - male chry.jpg Pseudagrion microcephalum by Sunny Joseph.jpg
Saffron-faced blue dart Pseudagrion rubricepsssp.ceylonicum* Pseudagrion rubriceps male-Kadavoor-2015-08-20-001.jpg Saffron-faced Blue Dart (Pseudagrion rubriceps)- Female W IMG 3613.jpg

Family: Platycnemididae - Threadtails

Common nameBinomialLocal subspeciesMaleFemale
Two-spotted threadtail Elattoneura oculata*
Jungle threadtail Elattoneura caesia* Elattoneura caesia 33483441.jpg
Dark-glittering threadtail Elattoneura centralis*
Smoky-winged threadtail Elattoneura leucostigma*


Red-striped threadtail Elattoneura tenax*
Stripe-headed threadtail Prodasineura sita*
Marsh dancer Onychargia atrocyana Onychargia atrocyana - Davidraju IMG 3023.jpg Onychargia atrocyana, Burdwan, West Bengal, India 13 09 2012.jpg
Yellow bush dart Copera marginipes Copera marginipes-Kadavoor-2016-11-11-001.jpg Copera marginipes-Kadavoor-2016-11-11-003.jpg

Family: Platystictidae - Shadowdamsels

Common nameBinomialLocal subspeciesMaleFemale
Alwis's shadowdamsel Ceylonosticta alwisi*
Ana Mia's shadowdamsel Ceylonosticta anamia*
Bine's shadowdamsel Ceylonosticta bine*
Ceylonosticta mirifica *
Ceylonosticta mojca *
Alwisi's shadowdamsel Ceylonosticta nancyae*
Rupasinghe's shadowdamsel Ceylonosticta rupasinghe*
Ceylonosticta venusta *
Drepanosticta adami *
Drepanosticta austeni *
Brinck's shadowdamsel Drepanosticta brincki*
Nobel shadowdamsel Drepanosticta digna*
Merry shadowdamsel Drepanosticta hilaris*
Drooping shadowdamsel Drepanosticta lankanensis*
Dark knob-tipped shadowdamsel Drepanosticta montana*
Nietner's shadowdamsel Drepanosticta nietneri*


Bordered knob-tipped shadowdamsel Drepanosticta submontana*
Blue-shouldered cornuted shadowdamsel Drepanosticta subtropica*
Dark-shouldered cornuted shadowdamsel Drepanosticta tropica*
Wall's shadowdamsel Drepanosticta walli*
Dark forestwraith Platysticta apicalis*
Blurry forestdamsel Platysticta maculata*
Platysticta secreta *
Platysticta serendibica *

Suborder: Anisoptera - Dragonflies.

A dragonfly is an insect belonging to the order Odonata, suborder Anisoptera (from Greek ἄνισος anisos "uneven" and πτερόν pteron, "wing", because the hindwing is broader than the forewing). Adult dragonflies are characterized by large multifaceted eyes, two pairs of strong transparent wings, sometimes with coloured patches and an elongated body. Dragonflies can be mistaken for the related group, damselflies (Zygoptera), which are similar in structure, though usually lighter in build; however, the wings of most dragonflies are held flat and away from the body, while damselflies hold the wings folded at rest, along or above the abdomen. Dragonflies are agile fliers, while damselflies have a weaker, fluttery flight. Many dragonflies have brilliant iridescent or metallic colours produced by structural coloration, making them conspicuous in flight. An adult dragonfly eye has nearly 24,000 ommatidia.

Dragonflies are predators, both in their aquatic larval stage, when they are known as nymphs or naiads, and as adults. Several years of their lives are spent as nymphs living in fresh water; the adults may be on the wing for just a few days or weeks. They are fast, agile fliers, sometimes migrating across oceans, and are often found near water.

There are 65 Dragonfly species can be found in Sri Lanka.

Family: Aeshnidae - Hawkers

Common nameBinomialLocal subspeciesMaleFemale
Donald's Hawker Anaciaeschna donaldi Anaciaeschna martini.jpg
Pale-spotted emperor Anax guttatus Anax guttatus by Nagabhushan Jyothi.jpg Anax guttatus female (13389411724).jpg
Magnificent emperor Anax immaculifrons Anax immaculifrons Blue Darner from Valparai IMG 8478 a.jpg Anax immaculifrons by Bala Chandran.jpg
Lesser green emperor Anax indicus Anax indicus by Arun V.jpg Anax indicus 5231.jpg
Black emperor Anax tristis Anax tristis 002176-1b.jpg
Vagrant emperor Anax ephippiger Hemianax ephippiger.jpg
Brown darner Gynacantha dravida Gynacantha-Kadavoor-2016-07-25-001.jpg Gynacantha-Kadavoor-2016-07-11-001.jpg
Gynacantha millardi Gynacantha millardi - Kadavoor - 001.jpg Gynacantha bayadera.jpg

Family: Gomphidae - Club-tail dragonflies

Common nameBinomialLocal subspeciesMaleFemale
Anisogomphus ceylonensis *
Sinuate clubtail Burmagomphus pyramidalisssp.sinuatus*
Cyclogomphus gynostylus *
Rivulet tiger Gomphidia pearsoni*
Lyrate grappletail Heliogomphus lyratus*

Heliogomphus lyratus 46804589.jpg

Nietner's grappletail Heliogomphus nietneri*
Wall's grappletail Heliogomphus walli*
Indian Common Clubtail Ictinogomphus rapax Indian common clubtail (Ictinogomphus rapax) male 3.jpg Ictinogomphus rapax (cropped).jpg
Keiser's forktail Macrogomphus annulatus*ssp.keiseri*


Sri Lankan Forktail Macrogomphus lankanensis* Sri Lanka Forktail Macrogomphus lankanensis.jpg
Microgomphus wijaya *
Sri Lanka sabretail Megalogomphus ceylonicus*
Brook hooktail Paragomphus henryi*
Paragomphus campestris *

Family: Macromiidae - Cruisers

Common nameBinomialLocal subspeciesMaleFemale
Common torrent hawk Epophthalmia vittatassp.cyanocephala* Epophthalmia vittata 3 by Suraj Naik.jpg Epophthalmia vittata (Common Torrent Hawk) female dorsal view.jpg
Macromia flinti *
Sri Lanka cruiser Macromia zeylanica*

Family: Libellulidae - Skimmers

Common nameBinomialLocal subspeciesMaleFemale
Asian pintail Acisoma panorpoidesssp.panorpoides Acisoma panorpoides-Kadavoor-2016-09-07-001.jpg Acisoma panorpoides-Kadavoor-2016-09-08-001.jpg
Scarlet marsh hawk Aethriamanta brevipennisssp.brevipennis Scarlet marsh hawk (Aethriamanta brevipennis) male.jpg Aethriamanta brevipennis 7678.jpg
Little Blue marsh hawk Brachydiplax sobrina Brachydiplax sobrina-Kadavoor-2016-06-27-005.jpg Little blue marsh hawk (Brachydiplax sobrina) female.jpg
Ditch jewel Brachythemis contaminata Brachythemis contaminata-male.jpg Brachythemis contaminata-female.jpg
Granite ghost Bradinopyga geminata Bradinopyga geminata at Kadavoor.jpg Bradinopyga geminata 6562.jpg
Scarlet skimmer Crocothemis serviliassp.servilia Crocothemis servilia male by kadavoor.jpg Crocothemis servilia female by kadavoor.jpg
Line forest-skimmer Cratilla lineatassp.calverti Cratilla lineata male 2 by kadavoor.jpg Cratilla lineata female by kadavoor.jpg
Black-tipped percher Diplacodes nebulosa Black Tipped Ground Skimmer (Diplacodes Nebulosa) male.jpg Black tipped ground skimmer female (36857328406).jpg
Chalky percher Diplacodes trivialis Diplacodes trivialis 05609.JPG Diplacodes trivialis, Burdwan, West Bengal, India 13 09 2012 5.jpg
Amber-winged marsh glider Hydrobasileus croceus Hydrobasileus croceus 00379.JPG Amber-winged Marsh Glider (Hydrobasileus croceus) female.,paanntt[?] prunt[?]. (43412314945).jpg
Fruhstorfer's junglewatcher Hylaeothemis fruhstorferissp.fruhstorferi*
Black marsh skimmer Indothemis carnatica Light-tipped demon (Indothemis carnatica) male Kanha.jpg Indothemis carnatica female by kadavoor.jpg
Restless demon Indothemis limbatassp.sita Indothemis limbata (cropped).jpg
Asiatic blood tail Lathrecista asiaticassp.asiatica Lathrecista asiatica 07056.JPG Lathrecista asiatica in Kerala.jpg
Lyriothemis defonsekai * Lyriothemis defonsekai.jpg
Coastal glider Macrodiplax cora Macrodiplax cora 4160.jpg Macrodiplax cora (5).jpg
Paddyfield parasol Neurothemis intermediassp.intermedia Neurothemis intermedia 05455.jpg Paddyfield parasol (Neurothemis intermedia) female Kanha.jpg
Pied paddy skimmer Neurothemis tullia Neurothemis tullia Male07798.jpg Pied paddy skimmer (Neurothemis tullia) female.jpg
Aggressive riverhawk Onychothemis tonkinensisssp.ceylanica
Spine-tufted skimmer Orthetrum chrysis Brown-backed Red Marsh Hawk Orthetrum chrysis Male 2 by Kadavoor.jpg Brown-backed red marsh hawk (Orthetrum chrysis) female Bardiya.jpg
Blue marsh hawk Orthetrum glaucum Orthetrum glaucum-Silent Valley-2016-08-14-002.jpg Orthetrum glaucum-Silent Valley-2016-08-14-003.jpg
Marsh skimmer Orthetrum luzonicum Orthetrum luzonicum-Kadavoor-2016-09-08-001.jpg Orthetrum luzonicum female in Kadavoor.jpg
Crimson-tailed marsh hawk Orthetrum pruinosumssp.neglectum Crimson-tailed marsh hawk (Orthetrum pruinosum neglectum) male Nepal.jpg Orthetrum pruinosum female by kadavoor.jpg
Slender skimmer Orthetrum sabinassp.sabina Green Marsh Hawk (Orthetrum sabina), Burdwan, West Bengal, India 22 09 2012.JPG Green marsh hawk (Orthetrum sabina) female Nepalgunj.jpg
Blue-tailed forest hawk Orthetrum triangularessp.triangulare Dragon Fly (Orthetrum triangulare) (7976467260).jpg Orthetrum triangulare by Rohit George.jpg
Globe skimmer Pantala flavescens Davidraju IMG 5948.jpg Pantala flavescens at Kadavoor.jpg
Yellow-tailed ashy skimmer Potamarcha congener Yellow-tailed Ashy Skimmer Potamarcha congener Male by kadavoor.jpg Yellow-tailed Ashy Skimmer Potamarcha congener juvenile male by kadavoor.jpg
Spine–legged redbolt Rhodothemis rufa Rufous Marsh Glider Rhodothemis rufa Male.jpg Rhodothemis rufa female by kadavoor.jpg
Sapphire flutterer Rhyothemis triangularis Rhyothemis triangularis-5822.jpg
Common picture wing Rhyothemis variegatassp.variegata Rhyothemis variegata - Davidraju IMG 4761.jpg Rhyothemis variegata female at Kadavoor.jpg


Red-veined darter Sympetrum fonscolombii Darter August 2007-20.jpg Darter August 2007-22 edit.jpg
Elf Tetrathemis yerburii* Tetrathemis yerburii.jpg
Coral-tailed cloudwing Tholymis tillarga Tholymis tillarga male by kadavoor 02.jpg Twister (Tholymis tillarga) female (16771052901).jpg
Crimson marsh glider Trithemis aurora Trithemis aurora male 2 by kadavoor.jpg Crimson marsh glider, female @ Thrippunithura 01.jpg
Black stream glider Trithemis festiva Black Stream Glider (Trithemis festiva)male (42030969054).jpg Black stream glider (Trithemis festiva) female. kaa[?]ttumpi. (36741083730).jpg


Dancing dropwing Trithemis pallidinervis Long-legged marsh glider (Trithemis pallidinervis) male Rajasthan.jpg LongLeggedMarshGlider 0915.JPG
Keyhole glider Tramea basilarisssp.burmeisteri Tramea basilaris,Red Marsh Trotter.jpg Keyhole Glider (Tramea basilaris) (11857165716).jpg
Black marsh trotter Tramea limbata Ferruginous glider (Tramea limbata) male 2.jpg Tramea limbata by Anubhav Agarwal.jpg
Greater crimson glider Urothemis signatassp.signata Greater Crimson Glider Urothemis signata Male by Kadavoor.jpg Greater Crimson Glider Urothemis signata Female by Kadavoor.jpg
Emerald cascader Zygonyx irisssp.ceylonicum Zygonyx iris-Thattekad-2015-09-12-001.jpg
Long-tailed duskdarter Zyxomma petiolatum Brown Dusk Hawk (Zyxomma petiolatum) (25368467958).jpg Zyxomma petiolatum by David Raju.jpg

See also

Related Research Articles

Odonata Order of insects that includes the dragonflies and damselflies

Odonata is an order of flying insects that includes the dragonflies and damselflies. Like most other flying insects, they evolved in the early Mesozoic era. Their prototypes, the giant dragonflies of the Carboniferous, 325 MYA, are no longer placed in the Odonata but included in the Protodonata or Meganisoptera.

Dragonfly Predatory winged insects

A dragonfly is a flying insect belonging to the order Odonata, infraorder Anisoptera. Adult dragonflies are characterized by a pair of large, multifaceted compound eyes, two pairs of strong, transparent wings, sometimes with coloured patches, and an elongated body. Dragonflies can be mistaken for the closely related damselflies, which make up the other odonatan infraorder (Zygoptera) and are similar in body plan though usually lighter in build; however, the wings of most dragonflies are held flat and away from the body, while damselflies hold their wings folded at rest, along or above the abdomen. Dragonflies are agile fliers, while damselflies have a weaker, fluttery flight. Many dragonflies have brilliant iridescent or metallic colours produced by structural colouration, making them conspicuous in flight. An adult dragonfly's compound eyes have nearly 24,000 ommatidia each.

Damselfly Suborder of insects

Damselflies are flying insects of the suborder Zygoptera in the order Odonata. They are similar to dragonflies, which constitute the other odonatan suborder, Anisoptera, but are smaller and have slimmer bodies. Most species fold the wings along the body when at rest, unlike dragonflies which hold the wings flat and away from the body. An ancient group, damselflies have existed since at least the Lower Permian, and are found on every continent except Antarctica.

Green darner Species of dragonfly

The green darner or common green darner, after its resemblance to a darning needle, is a species of dragonfly in the family Aeshnidae. One of the most common and abundant species throughout North America, it also ranges south to Panama. It is well known for its great migration distance from the northern United States south into Texas and Mexico. It also occurs in the Caribbean, Tahiti, and Asia from Japan to mainland China. It is the official insect for the state of Washington in the United States.

Blue-tailed damselfly Species of damselfly

The blue-tailed damselfly or common bluetail is a damselfly, belonging to the family Coenagrionidae.

Calopterygidae Family of damselflies

The Calopterygidae are a family of damselflies, in the suborder Zygoptera. They are commonly known as the broad-winged damselflies, demoiselles, or jewelwings. These rather large damselflies have wingspans of 50–80 mm, are often metallic-coloured, and can be differentiated from other damselflies by the broader connection between the wings and the body, as opposed to the abrupt narrowing seen in other damselfly families. The family contains some 150 species.

<i>Epiophlebia laidlawi</i> Species of dragonfly

The Himalayan relict dragonfly is one of four species of Epiprocta in the family Epiophlebiidae. They have at one time been classified as a suborder Anisozygoptera, considered as intermediate between the dragonflies and the damselflies, partly because the hind wings and fore wings are very similar in size and shape, and partly because the insect at rest holds them back over the body as damselflies do. These attributes now are known to be misleading however; the genus Epiophlebia shares a more recent ancestor with dragonflies and became separated from other Anisoptera in and around the uplifting Himalayas.

<i>Somatochlora</i> Genus of dragonflies

Somatochlora, or the striped emeralds, is a genus of dragonflies in the family Corduliidae with 42 described species found across the Northern Hemisphere.

<i>Megaloprepus caerulatus</i> Species of damselfly

Megaloprepus caerulatus is a damselfly of the Forest Giant family (Pseudostigmatidae), found in wet and moist forests in Central and South America. It has the greatest wingspan of any living damselfly or dragonfly, up to 19 centimeters (7.5 inches) in the largest males. Its large size and the markings on its wings make it a conspicuous species; a hovering Megaloprepus has been described as a "pulsating blue-and-white beacon".

<i>Austrolestes colensonis</i> Species of damselfly

Austrolestes colensonis, commonly known as the blue damselfly, is a species of damselfly of the family Lestidae. It is endemic to New Zealand and can commonly be found throughout the country, and at any time of the year. It is New Zealand's largest damselfly, and only blue odonate.

<i>Indolestes gracilis</i> Species of damselfly

Indolestes gracilis is a species of damselfly in the family Lestidae. It is known only from Sri Lanka, South India and Cambodia.

Drepanosticta subtropica is a species of damselfly in the family Platystictidae. It is endemic to Sri Lanka. The species was recorded for the first time after 83 years, along with three new species from Samanala Nature Reserve.

<i>Anax indicus</i> Species of dragonfly

Anax indicus is a species of dragonfly in the family Aeshnidae. It is found in India, Nepal, Pakistan, Sri Lanka, and Thailand.

<i>Bradinopyga geminata</i> Species of dragonfly

Bradinopyga geminata is a species of dragonfly in the family Libellulidae known commonly as the granite ghost. It is native to India, Sri Lanka and Thailand, where it is a common and widespread species.

Odonata are insects with an incomplete metamorphosis (hemimetabolous). The aquatic larva or nymph hatches from an egg, and develops through eight to seventeen instars before leaving the water and emerging as the winged adult or imago.

Ceylonosticta alwisi is a species of damselfly in the family Platystictidae. It is endemic to Sri Lanka, which was found recently from Samanala Nature Reserve, Ratnapura.

Alwisi's shadowdamsel is a species of damselfly in the family Platystictidae. It is endemic to Sri Lanka, which was found recently from Samanala Nature Reserve, Ratnapura.

Ceylonosticta rupasinghe is a species of damselfly in the family Platystictidae. It is endemic to Sri Lanka, which was found recently from Samanala Nature Reserve, Ratnapura.

<i>Indosticta deccanensis</i> Species of damselfly

Indosticta deccanensis, saffron reedtail is a damselfly species in the family Platystictidae. It is endemic to Western Ghats in India.

References

  1. Priyadarshana, Tharaka Sudesh; Wijewardhane, Ishara Harshajith; Herath, Bimal Eranga (29 November 2016). "Three new species of the genus Ceylonosticta Fraser, 1931 (Odonata: Zygoptera: Platystictidae) from Sri Lanka and the rediscovery of Ceylonosticta subtropica (Fraser, 1933". International Journal of Odonatology. 19 (4): 239–252. doi:10.1080/13887890.2016.1257443.
  2. "[Entomology • 2016] Taxonomy and Molecular Phylogeny of the Platystictidae (Insecta: Odonata) of Sri Lanka". Species New to Science. International Journal of Odonatology. 15 December 2016.