List of fauna of Batu Caves

The following species have been collected from the Dark Cave, Batu Caves, Selangor in Malaysia:

• Pheretima indica (Haplotaxida: Megascolecidae)
• Dichogaster sp. (Haplotaxida: Octochaetidae)
• Dichogaster bolavi (Haplotaxida: Octochaetidae)
• Sarax brachydactylus (Amblypygi: Tarantulidae)
• Damarchus cavernicolus (Araneae: Ctenizidae)
• Heteropoda robusta (Araneae: Heteropodidae)
• Liphistius batuensis (Araneae: Liphistiidae)
• Psiloderces crinitus (Araneae: Ochyroceratidae)
• Spermophora miser (Araneae: Pholcidae)
• Psechrus curvipalpus (Araneae: Psechridae)
• Scytodes magnus (Araneae: Scytodidae)
• Theriheridion rufipes (Araneae: Theridiidae)
• Uloborus spelaeus (Araneae: Uloboridae)
• Paracheyletia sp. (Eleutherengona: Cheyletidae)
• Ornithodorus bauensis (Ixodida: Argasidae)
• Belba sp. (Oribatida: Belbidae)
• Galumna sp. (Oribatida: Galumnidae)
• Vaghia sp. (Oribatida: Galumnidae)
• Scheloribates exuvium (Oribatida: Oribatulidae)
• Scheloribates sp. (Oribatida: Oribatulidae)
• Trombicula batui (Parasitengona: Trombiculidae)
• Trombicula insolli (Parasitengona: Trombiculidae)
• Cryptocheiridium lucifugum (Pseudoscorpionida: Ideoroncidae)
• Dhanus sumatranus (Pseudoscorpionida: Ideoroncidae)
• Dhanus doveri (Pseudoscorpionida: Ideoroncidae)
• Cunaxa setirostris (Trombidiformes: Cunaxidae)
• Scutigera decipiens (Scutigeromorpha: Cambalidae)
• Glyphiulus sp. (Sphaerotheriida: Cambalidae)
• Doratodesmus sp. (Sphaerotheriida: Cambalidae)
• Brachystomella contorta (Collembola: Brachystomellidae)
• Cunaxa spp. (Collembola: Cunaxidae)
• Willemia nadchatrami (Collembola: Neogastruridae)
• Aderus mcclurei (Coleoptera: Aderidae)
• Euglenes batuensis (Coleoptera: Aderidae)
• Euglenes cephalicus (Coleoptera: Aderidae)
• Euglenes malayanus (Coleoptera: Aderidae)
• Euglenes troglodytes (Coleoptera: Aderidae)
• Unidentified (Coleoptera: Carabidae)
• Unidentified (Coleoptera: Curculionidae)
• Aethriostoma undulata (Coleoptera: Dermestidae)
• Trinodes sp. (Coleoptera: Dermestidae)
• Cardiophorus carduelis (Coleoptera: Elateridae)
• Melanoxanthus near dohrni (Coleoptera: Elateridae)
• Platynuchus^{[ check spelling ]} sp. (Coleoptera: Elateridae)
• Unidentified (Coleoptera: Endomychidae)
• Platycladoxena near angulosa (Coleoptera: Erotylidae)
• Thallisellodes limbooliati (Coleoptera: Erotylidae)
• Cercyon gebieni (Coleoptera: Hydrophilidae)
• Dactylosternum abdominale (Coleoptera: Hydrophilidae)
• Lychnocrepis antricola (Coleoptera: Lampyridae)
• Unidentified (Coleoptera: Melyridae)
• Unidentified (Coleoptera: Nitidulidae)
• Unidentified (Coleoptera: Orthoperidae)
• Unidentified (Coleoptera: Pselaphidae)
• Trox costatus (Coleoptera: Scarabaeidae)
• Unidentified (Coleoptera: Scydmaenidae)
• Unidentified (Coleoptera: Silphidae)
• Unidentified (Coleoptera: Staphylinidae)
• Coeloecetes cavernicola (Coleoptera: Tenebrionidae)
• Chelisoches morio (Dermaptera: Forficulidae)
• Chelisoches brevipennis (Dermaptera: Forficulidae)
• Unidentified (Diptera: Cecidomyidae)
• Atrichopogon jacobsoni (Diptera: Ceratopogonidae)
• Culicoides huffi (Diptera: Ceratopogonidae)
• Culicoides peregrinus (Diptera: Ceratopogonidae)
• Culicoides prob. arakawai (Diptera: Ceratopogonidae)
• Forcipomyia spp. (Diptera: Ceratopogonidae)
• Stilobezzia sp. (Diptera: Ceratopogonidae)
• Eusmittia cavernae (Diptera: Chironomidae)
• Paratendipes inarmatus (Diptera: Chironomidae)
• Pentaneura batuensis (Diptera: Chironomidae)
• Podonomus sp. (Diptera: Chironomidae)
• Tricimba batucola (Diptera: Chloropidae)
• Chyromya prob. dubia (Diptera: Chyromyidae)
• Gymnochiromyia sp. (Diptera: Chyromyidae)
• Aedes albopictus (Diptera: Culicidae)
• Culex tritaeniorhynchus (Diptera: Culicidae)
• Uranotaenia sp. (Diptera: Culicidae)
• Condylostylus sp. (Diptera: Dolichopodidae)
• Drosophila ananassae (Diptera: Drosophilidae)
• Drosophila melanogaster (Diptera: Drosophilidae)
• Desmometopa spp. (Diptera: Milichiidae)
• Leptometopa mcclurei (Diptera: Milichiidae)
• Milichia sp. (Diptera: Milichiidae)
• Phyllomyza cavernae (Diptera: Milichiidae)
• Fannia leucosticta (Diptera: Muscidae)
• Ophyra chalcogaster (Diptera: Muscidae)
• Chetoneura cavernae (Diptera: Mycetophilidae)
• Eucampsipoda sundaicum (Diptera: Nycteribiidae)
• Nycteribosca prob. gigantea (Diptera: Nycteribiidae)
• Diploneura peregrina (Diptera: Phoridae)
• Brunettia sp. (Diptera: Psychodidae)
• Phlebotomus anodontis (Diptera: Psychodidae)
• Phlebotomus anondontis (Diptera: Psychodidae)
• Phlebotomus argentipes (Diptera: Psychodidae)
• Phlebotomus asperules (Diptera: Psychodidae)
• Phlebotomus lutea (Diptera: Psychodidae)
• Phlebotomus makati (Diptera: Psychodidae)
• Phlebotomus malayica (Diptera: Psychodidae)
• Phlebotomus savaiiensis (Diptera: Psychodidae)
• Phlebotomus stantion (Diptera: Psychodidae)
• Psychoda acanthostyla (Diptera: Psychodidae)
• Psychoda alternata (Diptera: Psychodidae)
• Psychoda aponesos (Diptera: Psychodidae)
• Psychoda harrisi (Diptera: Psychodidae)
• Psychoda malleola (Diptera: Psychodidae)
• Psychoda pellucida (Diptera: Psychodidae)
• Psychoda vagabunda (Diptera: Psychodidae)
• Sycorax malayensis (Diptera: Psychodidae)
• Telmatoscopus albipuntatus (Diptera: Psychodidae)
• Telmatoscopus kulas (Diptera: Psychodidae)
• Telmatoscopus mcclurei (Diptera: Psychodidae)
• Trichomyia batu (Diptera: Psychodidae)
• Trichomyia malaya (Diptera: Psychodidae)
• Unidentified (Diptera: Sarcophagidae)
• Bradysia flagellicornis (Diptera: Sciaridae)
• Bradysia leucocerca (Diptera: Sciaridae)
• Bradysia platytergum (Diptera: Sciaridae)
• Bradysia spp. (Diptera: Sciaridae)
• Bradysia leucocerca (Diptera: Sciaridae)
• Phorodonta malayana (Diptera: Sciaridae)
• Plastosciara near brevicalcarata (Diptera: Sciaridae)
• Soudekia sp. (Diptera: Sciaridae)
• Leptocera brevicostata (Diptera: Sphaeroceridae)
• Sargus metallinus (Diptera: Stratiomyidae)
• Helius cavernicolus (Diptera: Tipulidae)
• Cleon sp. (Ephemeroptera: Ephemeridae)
• Myiophanes fluitaria (Hemiptera: Reduviidae)
• Unidentified (Hemiptera: Cydnidae)
• Bagauda lucifigus (Hemiptera: Reduviidae)
• Reduvius gua (Hemiptera: Reduviidae)
• Fulvinus brevicornis (Hemiptera: Miridae)
• Apanteles carpatus (Hymenoptera: Braconidae)
• Apanteles sp. (Hymenoptera: Braconidae)
• Aulosaphes sp. (Hymenoptera: Braconidae)
• Epitranus lacteipennis (Hymenoptera: Chalcididae)
• Epitranus stantoni (Hymenoptera: Chalcididae)
• Bathroponera rufipes (Hymenoptera: Formicidae)
• Bathroponera tridentata (Hymenoptera: Formicidae)
• Camponotus sp. (Hymenoptera: Formicidae)
• Leptogenys diminuta (Hymenoptera: Formicidae)
• Monomorium pharaonis (Hymenoptera: Formicidae)
• Paratrechina longicornis (Hymenoptera: Formicidae)
• Pheidole javana (Hymenoptera: Formicidae)
• Ponera sp. (Hymenoptera: Formicidae)
• Pristomyrmex sp. (Hymenoptera: Formicidae)
• Tapinoma melanocephalum (Hymenoptera: Formicidae)
• Hypsicerca cavicola (Hymenoptera: Ichneumonidae)
• Hypsicerca fullawayi (Hymenoptera: Ichneumonidae)
• Trogaspidia sp. (Hymenoptera: Mutillidae)
• Opogona cerodelta (Lepidoptera: Lyonetidae)
• Attacus atlas (Lepidoptera: Saturniidae)
• Tinea antricola (Lepidoptera: Tineidae)
• Tinea palaechrysis (Lepidoptera: Tineidae)
• Neglurus vitripennis (Neuroptera: Myrmeleontidae)
• Diestrammena gravely (Orthoptera: Gryllacridae)
• Gryllotalpa fulvipes (Orthoptera: Gryllotalpidae)
• Mermecophilus dubius (Orthoptera: Gryllidae)
• Pycnoscelus striatus (Blattodea: Blaberidae)
• Liposcelis sp. (Psocoptera: Liposcelidae)
• Parasoa haploneura (Psocoptera: Lepidopsocidae)
• Psyllipsocus batuensis (Psocoptera: Psyllipsocidae)
• Ectopsocus maindroni (Psocoptera: Peripsocidae)
• Parasoa haploneura (Psocoptera: Forficulidae)
• Thaumapsylla breviceps orientalis (Siphonaptera: Ischnopsyllidae)
• Parabathynella malaya (Bathynellacea: Bathynellidae)
• Armadillo intermixtus (Isopoda: Armadillidiidae)
• Philoscia dobakholi (Isopoda: Oniscidae)
• Unidentified (Podocopida: Cyprididae)
• Chaerilus prob. celebensis (Scorpionida: Chactidae)
• Bufo asper (Anura: Bufonidae)
• Bufo melanostictus (Anura: Bufonidae)
• Callula pulchra (Anura: Microhylidae)
• Rana calconota (Squamata: Scincidae)
• Collocalia sp. (Apodiformes: Apodidae)
• Hirundo daurica (Passeriformes: Hirundinidae)
• Myophoneus flavirostris (Passeriformes: Turdidae)
• Taphozous melanopogon (Chiroptera: Emballonuridae)
• Eonycteris spelaea (Chiroptera: Pteropodidae)
• Hipposideros armiger debilis (Chiroptera: Rhinolophidae)
• Hipposideros bicolor (Chiroptera: Rhinolophidae)
• Hipposideros diadema vicarius (Chiroptera: Rhinolophidae)
• Hipposideros galiterus (Chiroptera: Rhinolophidae)
• Rhinolophus affinis superans (Chiroptera: Rhinolophidae)
• Rhinolophus luctus morio (Chiroptera: Rhinolophidae)
• Myotis mystacinus (Chiroptera: Vespertilionidae)
• Rattus rattus jalorensis (Rodentia: Muridae)
• Crocidura malayana (Soricomorpha: Soricidae)
• Elaphe taeniura (Squamata: Colubridae)
• Dryophiops rubescens (Squamata: Colubridae)
• Ahaetulla formosa (Squamata: Colubridae)
• Gecko marmorata (Squamata: Gekkonidae)
• Lygosoma scotophilum (Squamata: Scincidae)
• Paludomus buccula , Paludomus buccula var. minuta (Thiaridae)
• Opeas doveri (Stylommatophora: Stenogyridae)
• Opeas dimorpha (Stylommatophora: Stenogyridae)
• Achatina fulica (Pulmonata: Achatinidae)
• Dugesia sp. (Seriata: Scincidae)

References

• McClure, E.H., B.L. Lim and S.E. Winn. 1967. Fauna of Dark Cave, Batu Caves, Kuala Lumpur, Malaysia. Pacific Insects 9(3): 399-428.

Other references:

• Abraham, H.C. 1923. A new spider of the genus Liphistius. Journal of the Malayan Branch of the Royal Asiatic Society 1: 13-21.
• Annandale, N. and F.H. Gravely. 1914. The limestone caves of Burma and the Malay Peninsula, Part II: The fauna of the caves. Journal and Proceedings of the Asiatic Society of Bengal (ns) 9(10) for 1913: 402-423.
• Bristowe, W.S. 1933. The liphistiid spiders (chapters I-VIII, X). In Bristowe, W.S. and J. Millot, The liphistiid spiders, with an appendix on their internal anatomy. Proceedings of the Zoological Society of London 1932: 1016-1045, 1055-1057.
• Bristowe, W.S. 1952. The arachnid fauna of the Batu Caves in Malaya. Annals and Magazine of Natural History (ser. 12) 5: 697-707.
• Bullock, J.A. 1971. Fauna of Gua Anak Takun. Malayan Nature Journal 24: 95-97.
• Bullock, J.A. 1972. Cave Biology in Malaysia. Malayan Nature Journal 25: 135-141.
• Buxton, B.H. 1924. Notes on the internal anatomy of Liphistius batuensis Abr. Journal of the Malayan Branch of the Royal Asiatic Society 2(1): 85-86.
• Dittmar, K., M.L. Porter, L. Price, G. Svenson and M.F. Whitling. 2005. A Brief Survey of Invertebrates in Caves of Peninsular Malaysia. Malayan Nature Journal 57(2): 221-233.
• Dover, C. (ed.). 1929. Fauna of the Batu Caves, Selangor. Journal of the Federated Malay States Museums 14: 325-87.
• Dunn, F.L. 1965. Gua Anak Takun: Ecological Observations. Malayan Nature Journal 19(1): 75-87.
• Fage, L. 1929. Fauna of the Batu Caves, Selangor: x. Arachnida: Pedipalpi (Part) and Araneae. Journal of the Federated Malay States Museums 14: 356-64.
• Haupt, J. 2003. The Mesothelae - a monograph of an exceptional group of spiders (Araneae: Mesothelae): (Morphology, behavior, ecology, taxonomy, distribution and phylogeny). Zoologica 154: 1-102.
• Heynes-Wood, M. and C. Dover. 1929. Fauna of the Batu Caves, Selangor: iii. Topography. Journal of the Federated Malay States Museums 14: 329.
• Klingel, H. 1966. Beobachtungen an Lipistius batuensis Abr. (Araneae, Mesothelae). Zoologischer Anzeiger Supplementband 30: 246-253.
• Lim, T.T. 2006. A Visit to Gua Anak Takun. Pencinta Alam 2006(12): 14.
• Lim, T.W. and S.S. Yussof. 2009. Conservation status of Batu Caves Trapdoor Spider (Liphistius batuensis Abraham (Araneae, Mesothelae)): A preliminary survey. Malayan Nature Journal 61: 121-132.
• McClure, E.H. 1965. Microcosms of Batu Caves and a List of Species Collected at Batu Caves. Malayan Nature Journal 19(1): 68.
• McClure, E.H. 1995. Stories I like to Tell: An Autobiography. Elliot McClure [published by the author], Camarillo, CA. p. 197.
• Murphy, F. and J. Murphy. 2000. An Introduction to the Spiders of South East Asia. Kuala Lumpur: Malaysian Nature Society.
• Platnick, N.I. 2008. The World Spider Catalog, Version 9.0. American Museum of Natural History <research.amnh.org>.
• Platnick, N.I. and W.C. Sedgwick. 1984. A Revision of the Spider Genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1-31.
• Platnick, N.I. and W.J. Gertsch. 1976. The suborders of spiders: A cladistic analysis. American Museum Novitates 2607: 1-15.
• Price, L. 1996. The discovery of an ancient cave. Pencinta Alam 1996(4): 3.
• Ridley, H.N. 1899. Caves in the Malay Peninsula: Appendix. Report of the British Association for the Advancement of Science 1898: 580.
• Roewer, C.Fr. 1962. Einige arachniden aus den Batu-Caves in Malaya. Pacific Insects 4(2): 517-520.
• Schiödte, J.C. 1849. Om en afvigende Slaegt af Spindlernes Orden. Naturhistorisk Tidsskrift 2: 617-624.
• Soepadmo, E. and Ho Thian Hua. 1971. A Guide to Batu Caves. Kuala Lumpur: Malaysian Nature Society and Batu Caves Protection Association. p. 17.
• Yong, H.S. 1960. Field Biology: (d) Fauna of the Dark Cave (Batu Caves). Scientific Victorian 1960: 42-46.
• Yong, H.S. 1978. Living fossil spiders. Nature Malaysiana 3: 32-39.
• Yong, H.S. 1998. Spiders. The Encyclopedia of Malaysia 3: 94-95. Singapore: Archipelago Press.
• Yussof, S. 1985. The caves of Bukit Anak Takun. Nature Malaysiana 10(4): 4-9.
• Yussof, S. 1987. Gua Anak Takun Revisited. Malayan Naturalist 40(3&4): 12-16.
• Yussof, S. 1997. The natural and other histories of Batu Caves. Kuala Lumpur: Malaysian Nature Society.