Syzygium claviflorum | |
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Herbarium specimen of Szygium claviflorum | |
Scientific classification ![]() | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Rosids |
Order: | Myrtales |
Family: | Myrtaceae |
Tribe: | Syzygieae |
Genus: | Syzygium |
Species: | S. claviflorum |
Binomial name | |
Syzygium claviflorum | |
Synonyms [2] | |
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Syzygium claviflorum is a tree in the Myrtaceae family. It is native to northern and northeastern Australia and to tropical and subtropical Asia. It is used for timber, as fuel, as human and cattle food, and for dye. Stunted specimens can be found on the top of the plateau of Bokor National Park, Cambodia.
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This is an evergreen tree that grows to 10-20m tall in Southeast Asia, 3-15m tall in China. [3] [4] Trees in the north Queensland rainforests grow quickly to a large size, reaching some 180 to 270cm diameter. [5] The bark is often rather pale, greyish-white to greyish-brown. [6] The leaves have a depressed midrib and a grooved petiole on the upper surface, the blade is 6.5-13.2 by 2.9-5.8cm in size. Oil dots that are quite variable in size can be seen with a hand-lens. There are about 16-37 lateral veins on each side of the midrib.
The inflorescence bracts are deciduous, and absent at full flower opening, anthesis. In the flowers the calyx tube, hypanthium, and pedicel are around 10-20mm long together, the calyx tube is some 2-5mm in diameter, there are 4-5 rounded, small and inconspicuous calyx lobes, rarely greater than 0.5mm in length. There are 6-8 orbicular petals, when fully grown around 2-3mm in diameter, more or less cohering (largely as the apices of all petals fold into a cylinder at the centre between the staminal filaments). There are up to 20-30 oil dots on the petals. The outer staminal filaments are some 3-8mm long. Anthers are roughly 0.8 by 0.5mm in size, with a terminal gland, reasonably conspicuous. Each locule has about 9-16 ovules. The rather stout style is some 3-8mm long, the same or slightly longer than the stamens.
The edible fruit is of variable shape, from cylindrical to turbinate to pyriform. It is often galled and irregular, it is excavated at the apex and when mature 11-14 by 9-10mm in size. The calyx lobes are not obvious. The pericarp is succulent. Cells radiate from the endocarp to the exocarp. Seeds are about 7-8 by 7-8mm, with the testa apparently absent. The seedlings have about 3 pairs of reduced leaves, cataphylls, before the first pair of true leave appear, these cataphylls are some present between the leaves at later stages. At the tenth leaf stage, conspicuously quadrangled or shortly winged twigs appear, with a lanceolate to ovate leaf, whose apex is acuminate to acute, and a cuneate base. Oil dots are numerous, just visible to the naked eye, clearly visible with a hand-lens. Seeds germinate between 38-95 days.
The wood specific gravity is some 0.72-0.88, [6] wood density some 0.607g/cm3. [7]
Flowering occurs from January to March, June and November in Cambodia and Vietname, while fruiting is in March and April, June and August. [8] In Zhōngguó/China the tree flowers in March and April, fruiting in May and June. The specimens identified in Kerala, India flower and fruit from March to May, [9] while those in the Sikkim Himalaya fruit from August to October. [10]
Discussing the Papuasian subgenus Perkion, Craven distinguishes S. claviflorum in having the bark on the inflorescence branchlet of granular-papillate texture and having a flat staminal disc. [11] Traits used to distinguish S. claviflorum from other Szygium species in Cambodia and Vietnam are the large leaf blade (6-22 by 1.5-7.5cm); and the long petiole (0.3-0.6cm). [8]
This species is in the subgenus Perikion, for which it is the type species. [12] At a lower level it is in a clade with siblings Syzygium apodophyllum , S. corynanthum and Syzygium canicortex .
The tree grows on the Australian continent and across Wallacea, Southeast Asia to India and up to southern China. [2] [6] [4] [1] Countries and regions where the plant is native include: Australia (central- and north-eastern Queensland and Cape York; Yermalner/Melville Island, Bathurst Island, Northern Territory); Papua Niugini (eastern New Guinea); Indonesia (West Papua, Aru Islands, Sulawesi, Kalimantan, Jawa); Philippines; Malaysia (Sabah, Sarawak, Peninsular Malaysia); Thailand; Cambodia; Vietnam; Zhōngguó/China (Hainan, Yunnan); Laos; Myanmar; Bangladesh; India (Andaman Islands, Assam, Meghalaya, Sikkim); Bhutan; Eastern Himalaya; Sri Lanka. The species has been reported from southern India, in the southern Western Ghats in the state of Kerala. [9]
In Australia, the tree grows in well developed gallery forest and in lowland and upland rainforest on a variety of sites. [6] It occurs from sea level to 800m altitude. In Southeast Asia it primarily occurs in dense lowland forests. [3] In Zhōngguó/China the tree is found in dense or open angiosperm evergreen forest in valley and in the hills, with an elevation range of below 100m to 1300m. [4]
In North Queensland rainforests the taxa grows best on basic volcanic/basalt derived soils. [5]
Growing in New Guinea and the Aru Islands, the species occurs in a wide variety of communities: oak forest, rainforest, swamp forest, mangrove edges, riparian forest edges, riverine forest, lowland hill forest, tall open forest, forest on low stony hills, and oak-beech forest; at elevations from sea level to 2750m. [11]
The populations in Cambodia and Vietnam are found in lowland to montane forests, including primary, secondary and beach forest, up to 1050m elevation. [8]
On top of the plateau of Bokor National Park (Kampot Province, Cambodia, at some 1045m.a.s.l., this species grows as a shrub or small tree (2-4m tall) amongst a sclerophyllous stunted forest community on rocky soils. [13] [14] It is also reported as common in open bogs at between 930 and 1043m.
In the Chemunji Hills, Kerala it is common along streams in evergreen forest, while in Ponmudi, Kerala, it is common in exposed rocky areas of evergreen forests. [9] At both localities associated species are Polyalthia malabarica , Syzygium gardneri , S. lanceolatum , S. munroi , Elaeocarpus serratus var. weibelii, and Aporosa acuminata .
The "Least Concern" rating by the IUCN (see infobox above) is a result of the species having a large population, a very wide distribution and no current or future major threats have been identified. [1] However the IUCN notes that there is "[c]ontinuing decline in area, extent and/or quality of habitat." However in the Northern Territory, Australia, the plant is listed by the government as "Near Threatened", as the distribution of the tree is limited to Yermalner/Melville Island and Bathurst Island. [15] [16]
The wood is a useful structural timber, and in Australia is marketed under the name grey satinash. [6] People who live in the Bung Khong Long Non-Hunting Area (Nong Khai Province, Thailand) use the bark for dyeing nets (to enhance their strength) and the fruit as cattle feed. [17] In Cambodia the fruit are eaten and the wood is good firewood for heating. [3] In the Chittagong region of Myanmar and in Kerala and the Sikkim Himalaya, the fruit are eaten locally. [9] [10]
The species was described in 1841 by the German physician and botanist Ernst Gottlieb von Steudel (1783-1841), whose main work was on grasses. [18] He published the description in the second edition of his work Nomenclator botanicus (1840-1). [19] He was building on others work. Firstly that of Nathaniel Wallich (1786-1854), the Danmark-born botanist who worked in India, who did not quite validly describe the taxa. Before that, William Roxburgh (1751-1815), a Scottish surgeon and botanist who worked in India, had described the basionym taxa, that which this present accepted taxa is based: Eugenia claviflora Roxb. This was described in 1832 in Roxburgh's posthumously published work Flora Indica; or, Descriptions of Indian Plants.
Syzygium is a genus of flowering plants that belongs to the myrtle family, Myrtaceae. The genus comprises about 1200 species, and has a native range that extends from Africa and Madagascar through southern Asia east through the Pacific. Its highest levels of diversity occur from Malaysia to northeastern Australia, where many species are very poorly known and many more have not been described taxonomically. One indication of this diversity is in leaf size, ranging from as little as a half inch to as great as 4 ft 11 inches by sixteen inches in Syzygium acre of New Caledonia.
Syzygium smithii is a summer-flowering, winter-fruiting evergreen tree, native to Australia and belonging to the myrtle family Myrtaceae. It shares the common name "lilly pilly" with several other plants. It is planted as shrubs or hedgerows, and features: rough, woody bark; cream and green smooth, waxy leaves; flushes of pink new growth; and white to maroon edible berries. Unpruned, it will grow about 3–5 m (9.8–16.4 ft) tall in the garden.
Aglaia spectabilis is a species of tree in the family Meliaceae, found from the Santa Cruz Islands in the southwest Pacific to Queensland (Australia), Southeast Asia, Yunnan (Zhōngguó/China) and the Indian subcontinent. It grows from a 1m shrub to an emergent 40m tall tree, depending on the habitat. Its wood is commercially exploited as timber, but otherwise is of poor quality with limited use. The fruit are eaten, and used in folk medicine. The seeds are large in comparison to other plants, and a major source of dispersal of the species are hornbills eating the fruit, flying away from the tree and regurgitating the seeds.It is also found in Assam, India
Syzygium densiflorum is a species of evergreen tree in the family Myrtaceae. It is endemic to the Western Ghats mountains, India. The species is categorised as Vulnerable in the IUCN Red List.
Syzygium fibrosum is a species of rainforest trees native to monsoon forests of Indonesia, Papua New Guinea, and Australia. Common names include small red apple, (Bamaga) fibrous satinash and apricot satinash.
Syzygium hemilamprum, commonly known as the broad-leaved lilly pilly, blush satinash, cassowary gum, Eungella gum, and treated as Acmena hemilampra in New South Wales and Queensland, is a species of flowering plant in the family Myrtaceae and is native to New South Wales, Queensland and the Northern Territory. It is a rainforest tree with broadly lance-shaped to elliptic leaves, panicles of white flowers and more or less spherical white fruit.
Syzygium papyraceum, known as the paperbark satinash, is a rainforest tree of northeastern Queensland, Australia. It can grow to 35 m (115 ft) tall and 90 cm (35 in) in diameter, with papery red-brown bark. Leaves are simple, arranged in opposite pairs, and measure up to 10 cm (3.9 in) long and 5 cm (2.0 in) wide. Attractive pink or mauve flowers are followed by purple fruit. It was first described in 1983 by the Australian botanist Bernard Hyland.
Elaeocarpus lanceifolius is a tree species in the family Elaeocarpaceae. It is found across tropical Asia from Thailand to Yunnan to Nepal to Karnataka, India. It is used for its wood, fruit, and nuts.
Syzygium nervosum is a species of tree native to tropical Asia and Australia, extending as far north as Guangdong and Guangxi in China and as far south as the Northern Territory of Australia. It was previously known as Cleistocalyx operculatus and also known as C. nervosum(DC.) Kosterm., and Eugenia operculataRoxb., 1832. It is a medium-sized tree of about 10 meters in height with pale brown bark and dull green leaves.
Syzygium cormiflorum, commonly known as the bumpy satinash, is a species of Syzygium tree endemic to Queensland in northeastern Australia.
Syzygium alliiligneum, commonly known as onionwood, Mission Beach satinash or bark in the wood is a species of plant in the family Myrtaceae. It is endemic to a small part of north eastern Queensland.
Syzygium forte, commonly known as flaky-barked satinash, white apple or brown satinash, is a tree in the family Myrtaceae native to New Guinea and northern Australia.
Syzygium canicortex, commonly known as yellow satinash, is a tree in the family Myrtaceae native to Queensland, Australia, first described in 1983.
Syzygium apodophyllum is a tree in the Myrtaceae family endemic to north Queensland. The fruit is edible. It is a host for the exotic plant-pathogen fungus Austropuccinia psidii, which is causing a lot of damage to vegetation communities and economic plants.
Memecylon lilacinum is a tree species in the Melastomataceae family. It is usually an understorey species in closed forests. It is native to an area of tropical Asia, from Jawa to Philippines to Vietnam and the Andaman Islands and Myanmar. It is a food plant for the macaque Macaca facsicularis and a bee in the Megachilidae family.
Syzygium branderhorstii, commonly known as the Lockerbie satinash, is a small tree in the family Myrtaceae found in New Guinea, the Bismarck Archipelago, Solomon Islands, Santa Cruz Islands, and northern Queensland, Australia. It is cauliflorous, producing large inflorescences from the trunk. The fruits are eaten by brush turkeys.
Syzygium sayeri, commonly known as pink satinash, is a plant in the family Myrtaceae which is native to northeastern Queensland, Australia, and New Guinea.
Syzygium puberulum, commonly known as white satinash or downy satinash, is a plant in the family Myrtaceae which is native to rainforests of Cape York Peninsula, Queensland, and Papua New Guinea. It was first described in 1942.
Syzygium graveolens, commonly known as cassowary satinash, is a plant in the eucalyptus family Myrtaceae found only in the Wet Tropics bioregion of Queensland, Australia.
Syzygium maraca is a plant in the clove family Myrtaceae found only in the Wet Tropics bioregion of Queensland, Australia. It was first described in 2005 and has been given the conservation status of least concern. It has an affinity with Syzygium alliiligneum.
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