Tamisiocaris Temporal range: | |
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Movement of a frontal appendage | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | † Dinocaridida |
Order: | † Radiodonta |
Family: | † Tamisiocarididae |
Genus: | † Tamisiocaris Daley & Peel, 2010 |
Species: | †T. borealis |
Binomial name | |
†Tamisiocaris borealis Daley & Peel, 2010 | |
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The Cambrian explosion |
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Tamisiocaris (from Latin tamisium, sieve, and Greek karis, crab, shrimp) is a radiodont genus initially only known from frontal appendages from the Cambrian Sirius Passet lagerstatte in northern Greenland. It was initially described initially in 2010. [1] Further study in 2014 revealed that the frontal appendages were segmented and bore densely packed auxiliary spines, which were adapted to suspension feeding in a manner analogous to modern baleen whales. [2] It is assigned to the family Tamisiocarididae, and is measured about 22.8–33.6 cm (0.75–1.10 ft) long. [3]
Tamisiocaris in cladogram after Vinther et al., 2014. [2]
Radiodonta |
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Lobopodians are members of the informal group Lobopodia, or the formally erected phylum Lobopoda Cavalier-Smith (1998). They are panarthropods with stubby legs called lobopods, a term which may also be used as a common name of this group as well. While the definition of lobopodians may differ between literatures, it usually refers to a group of soft-bodied, marine worm-like fossil panarthropods such as Aysheaia and Hallucigenia.
Dinocaridida is a proposed fossil taxon of basal arthropods, which flourished during the Cambrian period and survived up to the Early to Middle Ordovician. The putative Early Devonian member Schinderhannes bartelsi has been considered questionable by some researchers.
Sirius Passet is a Cambrian Lagerstätte in Peary Land, Greenland. The Sirius Passet Lagerstätte was named after the Sirius sledge patrol that operates in North Greenland. It comprises six places in Nansen Land, on the east shore of J.P. Koch Fjord in the far north of Greenland. It was discovered in 1984 by A. Higgins of the Geological Survey of Greenland. A preliminary account was published by Simon Conway Morris and others in 1987 and expeditions led by J. S. Peel and Conway Morris have returned to the site several times between 1989 and the present. A field collection of perhaps 10,000 fossil specimens has been amassed. It is a part of the Buen Formation.
Anomalocaris is an extinct genus of radiodont, an order of early-diverging stem-group arthropods.
Peytoia is a genus of hurdiid radiodont, an early diverging order of stem-group arthropods, that lived in the Cambrian period, containing two species, Peytoia nathorsti from the Miaolingian of Canada and Peytoia infercambriensis from Poland, dating to Cambrian Stage 3. Its two frontal appendages had long bristle-like spines, it had no fan tail, and its short stalked eyes were behind its large head.
Anomalocarididae is an extinct family of Cambrian radiodonts, a group of stem-group arthropods.
Kerygmachela kierkegaardi is a kerygmachelid gilled lobopodian from the Cambrian Stage 3 aged Sirius Passet Lagerstätte in northern Greenland. Its anatomy strongly suggests that it, along with its relative Pambdelurion whittingtoni, was a close relative of radiodont and euarthropods. The generic name "Kerygmachela" derives from the Greek words Kerygma (proclamation) and Chela (claw), in reference to the flamboyant frontal appendages. The specific name, "kierkegaardi" honors Danish philosopher Søren Kierkegaard.
Pambdelurion is an extinct genus of panarthropod from the Cambrian aged Sirius Passet site in northern Greenland. Like the morphologically similar Kerygmachela from the same locality, Pambdelurion is thought to be closely related to arthropods, combining characteristics of "lobopodians" with those of primitive arthropods.
Schinderhannes bartelsi is a species of stem-group arthropod, possibly a hurdiid radiodont (anomalocaridid), known from one specimen from the Lower Devonian Hunsrück Slates. Its discovery was astonishing because the latest definitive radiodonts were known only from the Early Ordovician, at least 66 million years earlier than this taxon. Although some phylogenetic analyses found support for its taxonomic placement as a hurdiid radiodont, other researchers have questioned these assignments.
Radiodonta is an extinct order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two originally refer to the family Anomalocarididae, which previously included all species of this order but is now restricted to only a few species. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and used for a variety of functions. Radiodonts included the earliest large predators known, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa Anomalocaris canadensis, Hurdia victoria, Peytoia nathorsti, Titanokorys gainessii, Cambroraster falcatus and Amplectobelua symbrachiata. The latest surviving members include the Aegirocassisinae from the Early Ordovician of Morocco. The putative Early Devonian member Schinderhannes bartelsi from Germany has been considered questionable by some researchers.
Caryosyntrips ("nutcracker") is an extinct genus of stem-arthropod which known from Canada, United States and Spain during the middle Cambrian.
Kiisortoqia soperi is an extinct species of arthropod from the Early Cambrian Sirius Passet Lagerstätte in Greenland. While it had a superficially trilobite-like bodyform, it also possessed large frontal appendages similar to those of radiodonts.
Tamisiocarididae is a family of radiodonts, extinct marine animals related to arthropods, that bore finely-spined appendages that were presumably used in filter-feeding. When first discovered, the clade was named Cetiocaridae after a speculative evolution artwork, Bearded Ceticaris by John Meszaros, that depicted a hypothetical filter-feeding radiodont at a time before any were known to exist. However, the family name was not valid according to the International Code of Zoological Nomenclature, as no real genus named "Cetiocaris" exists, and in 2019 it was formally replaced by the name Tamisiocarididae, after the only valid genus of the clade at the time. The family is only known from Series 2 of the Cambrian, unlike other radiodont families, which persisted longer into the Cambrian. All known species would have lived in tropical or subtropical waters, suggesting a preference for warmer waters.
Amplectobeluidae is a clade of Cambrian radiodonts. It currently includes five definitive genera, Amplectobelua, Lyrarapax, Ramskoeldia, Guanshancaris and a currently unnamed genus from the lower Cambrian aged Sirius Passet site in Greenland. There is also a potential fifth genus, Houcaris, but that genus has become problematic in terms of its taxonomic placement.
Hurdiidae is an extinct cosmopolitan family of radiodonts, a group of stem-group arthropods, which lived during the Paleozoic Era. It is the most long-lived radiodont clade, with definitive members lasting from the Cambrian to the earliest Ordovician period. The putative Early Devonian member Schinderhannes bartelsi has been considered questionable by some researchers.
Houcaris is a possibly paraphyletic radiodont genus, tentatively assigned to either Amplectobeluidae, Anomalocarididae or Tamisiocarididae, known from Cambrian Series 2 of China and the United States. The type species is Houcaris saron which was originally described as a species of the related genus Anomalocaris. Other possible species include H. magnabasis and H. consimilis. The genus Houcaris was established for the two species in 2021 and honors Hou Xianguang, who had discovered and named the type species Anomalocaris saron in 1995 along with his colleagues Jan Bergström and Per E. Ahlberg.
Pahvantia is an extinct genus of hurdiid radiodont from the Cambrian. It is known by a single species, Pahvantia hastata, described from Wheeler Shale and Marjum Formation in Utah. Although it was once considered as filter feeder using large number of putative setae, this structures are later considered as misidentification of trunk materials.
Pseudoangustidontus is a genus of hurdiid (peytoiid) radiodont from the Lower Ordovician of Morocco. This genus is known from two described species, P. duplospineus and P. izidigua, with some specimens that are hard to determine which species to belong to. This animal is only known from the Fezouata Formation, a fossil site in Morocco that is of Lagerstätte status, meaning that the fossils from this site are exceptionally preserved. Because of its partial remains, its classification was debated, but with more complete fossils it is identified as radiodont frontal appendage.
Aegirocassisinae is a subfamily of radiodonts from the lower Paleozoic era. It belongs to the larger hurdiidae (peytoiid) family, which were the most diverse and long lasting radiodonts. The members of this subfamily are restricted to the Lower Ordovician-aged Fezouata Formation of Morocco. Currently only two genera are included: Aegirocassis and Pseudoangustidontus. These two genera possess large Baleen-like auxiliary spines on their frontal appendages, which suggests a suspension feeding lifestyle for the group. These radiodonts are some of the few known from sediments beyond the Cambrian period. This subfamily shows that following the Great Ordovician Biodiversification Event, which saw a rise in the plankton population in the worlds oceans, suspension feeding became more common in radiodonts then other feeding styles. It also seems that due to the evolution of new predators, like large nautiloid cephalopods, and other arthropod groups like the eurypterids, the radiodonts evolved suspension feeding lifestyles in order to minimize competition for food.
Bushizheia yangi is an extinct species of arthropod known from a single specimen found in the Early Cambrian Maotianshan Shales Lagerstätte in China. While it had a superficially trilobite-like bodyform, it also possessed large frontal appendages similar to those of radiodonts. It is morphologically similar to Kiisortoqia to which it is probably closely related.