Titanohyrax Temporal range: | |
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Teeth | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Hyracoidea |
Family: | † Titanohyracidae |
Genus: | † Titanohyrax Matsumoto , 1922 [1] |
Type species | |
Titanohyrax andrewsi | |
Species | |
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Titanohyrax is an extinct genus of large to very large hyrax from the Eocene and Oligocene. Specimens have been discovered in modern-day Algeria, Tunisia, Egypt and Libya. Some species, like T. ultimus, are estimated to be as large as the modern rhinoceros. Titanohyrax species are still poorly known due to their rarity in the fossil record.
Titanohyrax is unusual among the numerous Paleogene hyracoids by its lophoselenodont teeth (having teeth that are lophodont and selenodont), fully molariform premolars, and relatively high-crowned cheek teeth. This suggests the genus had a folivorous diet. [2]
The genus was first described by in 1922 for the species T. ultimus from the early Oligocene of the Jebel Qatrani Formation, Fayum Depression, Egypt. [3] The author described it as an “extremely gigantic species, being the largest of all the hyracoids hitherto known” – estimates of body mass range from 600 kg (1,300 lb) to 1,300 kg (2,900 lb). [4] T. tantulus is the smallest Titanohyrax species known, with a body mass of around 23 kg (51 lb). [4]
Moeritherium is an extinct genus of basal proboscideans from the Eocene of North and West Africa. It was related to elephants and, more distantly, to sea cows and hyraxes.
Aegyptopithecus is an early fossil catarrhine that predates the divergence between hominoids (apes) and cercopithecids. It is known from a single species, Aegyptopithecus zeuxis, which lived around 38-29.5 million years ago in the early part of the Oligocene epoch. It likely resembled modern-day New World monkeys, and was about the same size as a modern howler monkey, which is about 56 to 92 cm long. Aegyptopithecus fossils have been found in the Jebel Qatrani Formation of modern-day Egypt. Aegyptopithecus is believed to be a stem-catarrhine, a crucial link between Eocene and Miocene fossils.
Arsinoitherium is an extinct genus of paenungulate mammals belonging to the extinct order Embrithopoda. It is related to elephants, sirenians, and hyraxes. Arsinoitheres were superficially rhinoceros-like herbivores that lived during the Late Eocene and the Early Oligocene of North Africa from 36 to 30 million years ago, in areas of tropical rainforest and at the margin of mangrove swamps. A species described in 2004, A. giganteum, lived in Ethiopia about 27 million years ago.
Paraceratherium is an extinct genus of hornless rhinocerotoids belonging to the family Paraceratheriidae. It is one of the largest terrestrial mammals that has ever existed and lived from the early to late Oligocene epoch. The first fossils were discovered in what is now Pakistan, and remains have been found across Eurasia between China and the Balkans. Paraceratherium means "near the hornless beast", in reference to Aceratherium, the genus in which the type species P. bugtiense was originally placed.
Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring 5 metres (16 ft) long and weighing 1–2.2 metric tons. Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand and Western Sahara.
Megistotherium is an extinct genus of hyaenodont belonging to the family Hyainailouridae that lived in Africa.
Coryphodon is an extinct genus of pantodonts of the family Coryphodontidae.
Barytherium is a genus of an extinct family (Barytheriidae) of primitive proboscideans that lived during the late Eocene and early Oligocene in North Africa. The type species is Barytherium grave, found at the beginning of the 20th century in Fayum, Egypt. Since then, more complete specimens have been found at Dor el Talha, Libya. More fossils were also discovered in 2011 in the Aidum area in Dhofar by Oman's Ministry of Heritage and Culture, which was named Barytherium omansi.
Hoplophoneus is an extinct genus of the family Nimravidae, endemic to North America during the Late Eocene to Early Oligocene epochs, existing for approximately 6 million years.
Ptolemaiida is a taxon of wolf-sized afrothere mammals that lived in northern and eastern Africa during the Paleogene. The oldest fossils are from the latest Eocene strata of the Jebel Qatrani Formation, near the Fayum oasis in Egypt. A tooth is known from an Oligocene-aged stratum in Angola, and Miocene specimens are known from Kenya and Uganda.
Witwatia is an extinct genus of giant bat that contained two species which lived in the Al Fayyum in Egypt during the late Eocene and one species which lived in Tunisia during the early Eocene. It is known from a lower jaw and teeth. Three species have been named: the type species W. schlosseri, W. eremicus and W. sigei.
The Jebel Qatrani Formation is a geologic formation located in the Faiyum Governorate of central Egypt. It is exposed between the Jebel Qatrani escarpment and the Qasr el Sagha escarpment, north of Birket Qarun lake near Faiyum. The formation conformably overlies the Qasr el Sagha Formation and is topped by the Widan el Faras Basalt. The age of the formation has been subject to debate, but the most recent research indicates that it covers both the latest parts of the Eocene and the Early Oligocene, spanning over the boundary between these two time periods.
Pterodon is an extinct genus of hyaenodont in the family Hyainailouridae, containing five species. The type species Pterodon dasyuroides is known exclusively from the late Eocene to the earliest Oligocene of western Europe. The genus was first erected by the French zoologist Henri Marie Ducrotay de Blainville in 1839, who said that Georges Cuvier presented one of its fossils to a conference in 1828 but died before he could make a formal description of it. It was the second hyaenodont genus with taxonomic validity after Hyaenodon, but this resulted in taxonomic confusion over the validities of the two genera by other taxonomists. Although the taxonomic status of Pterodon was revised during the late 19th and early 20th centuries, it became a wastebasket taxon for other hyaenodont species found in Africa and Asia. Today, only the type species is recognized as belonging to the genus while four others are pending reassessment to other genera.
Hyraxes, also called dassies, are small, stout, herbivorous mammals in the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Modern hyraxes are typically between 30 and 70 cm in length and weigh between 2 and 5 kg. They are superficially similar to marmots, or over-large pikas, but are much more closely related to elephants and sea cows. Hyraxes have a life span from 9 to 14 years. Both types of "rock" hyrax live on rock outcrops, including cliffs in Ethiopia and isolated granite outcrops called koppies in southern Africa.
Megalohyrax is an extinct hyrax-grouped genus of herbivorous mammal that lived during the Oligocene, about 33-30 million years ago. Its fossils have been found in Africa and in Asia Minor.
Propliopithecoidea is a superfamily of catarrhine primates that inhabited Africa and the Arabian Peninsula during the Early Oligocene about 32 to 29 million years ago. Fossils have been found in Egypt, Oman and Angola. They are one of the earliest known families of catarrhines. They have a number of features in common with extant catarrhines, but also features that are primitive and not found in later catarrhine families.
Phiomicetus is a genus of protocetid whale that lived between 43 and 42 million years ago during the Lutetian period in what is now Egypt. It had powerful jaws and large teeth that would have allowed it to hunt and tear large prey.
Sparnotheriodon is an extinct genus of sparnotheriodontid litoptern that lived during the Middle Eocene of what is now Argentina, leaving fossils in the Sarmiento Formation.
Mixtotherium is an extinct genus of Palaeogene artiodactyls belonging to the monotypic family Mixtotheriidae. Known informally as mixtotheriids or mixtotheres, these artiodactyls were endemic to western Europe and occurred from the middle to late Eocene. The genus and type species were both first established by the French naturalist Henri Filhol in 1880. Several species are well known by good skull fossils, which were informative enough to allow for classifications of the species to their own family. The Mixtotheriidae, first recognized by Helga Sharpe Pearson in 1927, is currently known by 7 valid species, although M. priscum is thought by several authors to be synonymous with M. gresslyi. The affinities of the Mixtotheriidae in relation to other artiodactyl families is uncertain, but it is currently thought to have been related to the Cainotherioidea and Anoplotheriidae.
The Fayyum is a region and an important fossil Lagerstätte in northern Egypt. The region comprises the Fayyum Basin, which is intensively used for agriculture, and adjoining areas; the important areas of discovery are mostly north and west of Lake Qarun. The Wadi al-Hitan, known for its numerous whale fossils and since 2005 UNESCO World Natural Heritage Site, forms the south-western end. The deposits of the Fayyum belong to various geological formations. They are mainly composed of limestone, siltstone and sandstone. The lower sections consist of marine sediments, while the upper, continental sediments were formed in a coastal landscape. The formation period ranges from the Middle to the Upper Eocene to the Lower Oligocene, which corresponds to an age of around 41 to 28 million years ago. The entire sedimentary complex is overlain by basalt, which dates back to volcanic activity around 24 million years ago.