Amaranthus tuberculatus | |
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1913 illustration [1] | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Order: | Caryophyllales |
Family: | Amaranthaceae |
Genus: | Amaranthus |
Species: | A. tuberculatus |
Binomial name | |
Amaranthus tuberculatus (Moq.) J.D.Sauer | |
Synonyms [3] | |
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Amaranthus tuberculatus, commonly known as roughfruit amaranth, [4] rough-fruited water-hemp, [5] tall waterhemp, or common waterhemp, [6] [7] is a species of flowering plant. It is a summer annual broadleaf with a germination period that lasts several months. [8] Tall waterhemp has been reported as a weed in 40 of 50 U.S. states. [9]
A distinguishing characteristic of tall waterhemp that sets it apart from similar members of the genus Amaranthus is the lack of hair on its stems and leaves. This characteristic gives the plant a bright, glossy appearance.
The leaves of tall waterhemp tend to be long and narrow.
The stem is typically erect and slender and can be up to three feet long. The color of the stem is green or red. [8]
Tall waterhemp is a dioecious plant. The seedhead branches in the female are numerous, short, and smooth. The male seedhead branches are fewer, longer, and more slender than those of the female.
The species has terminal spike inflorescences and very short bracts with simple to highly branched flowers. Seed produced is reddish to black and less than 1/32 inch in diameter. [10]
Tall waterhemp is native to North America. It is believed to have originally had a range north of Missouri and Tennessee to the Great Lakes. [11] It is now found in 40 states but is most common in the Great Plains and Great Lakes regions. [12]
Tall waterhemp predominantly grows in wet habitats, such as ponds, marshes, lakes, creeks, and other riparian zones. It also thrives along roadways and railroads as well as agricultural fields. [11] It can grow in a variety of climates, as evidenced by its widespread range.
Tall waterhemp is a summer annual that produces a large number[ quantify ] of very small[ quantify ] seeds. It is considered an r-strategist. Emergence can span several months and often occurs later in the season than other annual weeds, allowing the weed to evade typical weed control strategies such as herbicide application and tillage. [8] One study observed 80% emergence not occurring until ten weeks after the initial emergence. [13] Extreme temperatures have little effect on seed viability. Germination occurs typically after soil temperature alternation, as this is required to break seed dormancy. [14] Waterhemp has been found to germinate in a wide range of soil and temperature conditions. [15] It has been found to germinate 17 years after seed set. [16]
While tall waterhemp cannot self-pollinate, due to having separate male and female plants, it does not require any vectors for pollination. [15] This allows for wind pollination over large distances, generating a large amount of genetic diversity. Another factor contributing to genetic diversity is the large amount of seed produced. Tall waterhemp in competition with soybean has been reported to produce from between 300,000 and 5,000,000 seeds per plant. [17] Tall waterhemp also has a rapid growth rate, 50–70% greater than other annual weeds. [18]
Interspecific hybrids of tall waterhemp and Amaranthus hybridus have been observed in experimental fields [19] but have not been observed in agronomic fields. [20]
In North America, tall waterhemp is considered a major weed of agricultural fields and other disturbed habitats. [11] The Southern Weed Science Society includes tall waterhemp on their list of weed species. [21] However, it is not listed on the federal noxious weed list or any state lists in the United States. [22] In Europe and other continents where the species has been introduced, naturalization is an infrequent occurrence. [11]
Because of the long germination window for tall waterhemp, a single herbicide application is unlikely to be an effective control strategy. [23] Michigan State University Extension recommends a preemergence application followed by one or more postemergence applications. [8] [23] Some populations of tall waterhemp have been reported resistant to acetolactate synthase inhibiting (ALS) herbicides [23] and the triazines, [23] with some individual weeds being resistant to both herbicide groups. [23] Resistance to acifluorfen and other diphenyl ether herbicides has been reported. [8] [15] Even more alarming is the emergence of waterhemp resistant to the latest generation of herbicides, HPPD inhibitors. [24] [23] Furthermore, waterhemp at a site in Nebraska was found to be resistant to 2,4-D, a phenoxy herbicide. [25] [23]
According to Bob Hartzler of Iowa State University, the most effective control of tall waterhemp is achieved by cultural practices that promote growth of the desired vegetation. [15]
Amaranthus is a cosmopolitan genus of annual or short-lived perennial plants collectively known as amaranths. Some amaranth species are cultivated as leaf vegetables, pseudocereals, and ornamental plants. Catkin-like cymes of densely packed flowers grow in summer or fall. Amaranth varies in flower, leaf, and stem color with a range of striking pigments from the spectrum of maroon to crimson and can grow longitudinally from 1 to 2.5 metres tall with a cylindrical, succulent, fibrous stem that is hollow with grooves and bracteoles when mature. There are approximately 75 species in the genus, 10 of which are dioecious and native to North America with the remaining 65 monoecious species endemic to every continent from tropical lowlands to the Himalayas. Members of this genus share many characteristics and uses with members of the closely related genus Celosia. Amaranth grain is collected from the genus. The leaves of some species are also eaten.
Herbicides, also commonly known as weed killers, are substances used to control undesired plants, also known as weeds. Selective herbicides control specific weed species while leaving the desired crop relatively unharmed, while non-selective herbicides can be used to clear waste ground, industrial and construction sites, railways and railway embankments as they kill all plant material with which they come into contact. Apart from selective/non-selective, other important distinctions include persistence, means of uptake, and mechanism of action. Historically, products such as common salt and other metal salts were used as herbicides, however, these have gradually fallen out of favor, and in some countries, a number of these are banned due to their persistence in soil, and toxicity and groundwater contamination concerns. Herbicides have also been used in warfare and conflict.
Senecio vulgaris, often known by the common names groundsel and old-man-in-the-spring, is a flowering plant in the family Asteraceae. It is an annual herb, native to the Palaearctic and widely naturalised as a ruderal species in suitable disturbed habitats worldwide.
Weed control is a type of pest control, which attempts to stop or reduce growth of weeds, especially noxious weeds, with the aim of reducing their competition with desired flora and fauna including domesticated plants and livestock, and in natural settings preventing non native species competing with native species.
Bromus tectorum, known as downy brome, drooping brome or cheatgrass, is a winter annual grass native to Europe, southwestern Asia, and northern Africa, but has become invasive in many other areas. It now is present in most of Europe, southern Russia, Japan, South Africa, Australia, New Zealand, Iceland, Greenland, North America and western Central Asia. In the eastern US B. tectorum is common along roadsides and as a crop weed, but usually does not dominate an ecosystem. It has become a dominant species in the Intermountain West and parts of Canada, and displays especially invasive behavior in the sagebrush steppe ecosystems where it has been listed as noxious weed. B. tectorum often enters the site in an area that has been disturbed, and then quickly expands into the surrounding area through its rapid growth and prolific seed production.
Onopordum acanthium is a flowering plant in the family Asteraceae. It is native to Europe and Western Asia from the Iberian Peninsula east to Kazakhstan, and north to central Scandinavia, and widely naturalised elsewhere, with especially large populations present in the United States and Australia. It is a vigorous biennial plant with coarse, spiny leaves and conspicuous spiny-winged stems.
Euphorbia heterophylla, also known under the common names of Mexican fireplant, painted euphorbia, Japanese poinsettia, paintedleaf, painted spurge and milkweed, is a plant belonging to the Euphorbiaceae or spurge family.
Centaurea solstitialis, the yellow star-thistle, is a species of thorny plant in the genus Centaurea, which is part of the family Asteraceae. A winter annual, it is native to the Mediterranean Basin region and invasive in many other places. It is also known as golden starthistle, yellow cockspur and St. Barnaby's thistle.
In agriculture, a living mulch is a cover crop interplanted or undersown with a main crop, and intended to serve the purposes of a mulch, such as weed suppression and regulation of soil temperature. Living mulches grow for a long time with the main crops, whereas cover crops are incorporated into the soil or killed with herbicides.
Abutilon theophrasti is an annual plant in the family Malvaceae that is native to southern Asia. Its specific epithet theophrasti commemorates the ancient Greek botanist-philosopher Theophrastus. Abutilon theophrasti is the type species of the genus Abutilon.
Amaranthus palmeri is a species of edible flowering plant in the amaranth genus. It has several common names, including carelessweed, dioecious amaranth, Palmer's amaranth, Palmer amaranth, and Palmer's pigweed.
Melilotus albus, known as honey clover, white melilot (UK), Bokhara clover (Australia), white sweetclover (US), and sweet clover, is a nitrogen-fixing legume in the family Fabaceae. Melilotus albus is considered a valuable honey plant and source of nectar and is often grown for forage. Its characteristic sweet odor, intensified by drying, is derived from coumarin.
Orobanche aegyptiaca, the Egyptian broomrape, is a plant which is an obligate holoparasite from the family Orobanchaceae with a complex lifecycle. This parasite is most common in the Middle East and has a wide host range including many economically important crops.
A weed is a plant considered undesirable in a particular situation, growing where it conflicts with human preferences, needs, or goals. Plants with characteristics that make them hazardous, aesthetically unappealing, difficult to control in managed environments, or otherwise unwanted in farm land, orchards, gardens, lawns, parks, recreational spaces, residential and industrial areas, may all be considered weeds. The concept of weeds is particularly significant in agriculture, where the presence of weeds in fields used to grow crops may cause major losses in yields. Invasive species, plants introduced to an environment where their presence negatively impacts the overall functioning and biodiversity of the ecosystem, may also sometimes be considered weeds.
Striga hermonthica, commonly known as purple witchweed or giant witchweed, is a hemiparasitic plant that belongs to the family Orobanchaceae. It is devastating to major crops such as sorghum and rice. In sub-Saharan Africa, apart from sorghum and rice, it also infests maize, pearl millet, and sugar cane.
Mesotrione is the ISO common name for an organic compound that is used as a selective herbicide, especially in maize. A synthetic inspired by the natural substance leptospermone, it inhibits the enzyme 4-hydroxyphenylpyruvate dioxygenase (HPPD) and is sold under brand names including Callisto and Tenacity. It was first marketed by Syngenta in 2001.
4-Hydroxyphenylpyruvate dioxygenase (HPPD) inhibitors are a class of herbicides that prevent growth in plants by blocking 4-Hydroxyphenylpyruvate dioxygenase, an enzyme in plants that breaks down the amino acid tyrosine into molecules that are then used by plants to create other molecules that plants need. This process of breakdown, or catabolism, and making new molecules from the results, or biosynthesis, is something all living things do. HPPD inhibitors were first brought to market in 1980, although their mechanism of action was not understood until the late 1990s. They were originally used primarily in Japan in rice production, but since the late 1990s have been used in Europe and North America for corn, soybeans, and cereals, and since the 2000s have become more important as weeds have become resistant to glyphosate and other herbicides. Genetically modified crops are under development that include resistance to HPPD inhibitors. There is a pharmaceutical drug on the market, nitisinone, that was originally under development as an herbicide as a member of this class, and is used to treat an orphan disease, type I tyrosinemia.
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Jonathan Gressel is an Israeli agricultural scientist and Professor Emeritus at the Weizmann Institute of Science in Rehovot, Israel. Gressel is a "strong proponent of using modern genetic techniques to improve agriculture" especially in third world and developing countries such as Africa. In 2010, Gressel received Israel's highest civilian award, the Israel Prize, for his work in agriculture
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