Ankylosaurines Temporal range: Cretaceous, | |
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Skeleton of Scolosaurus thronus , once referred to Euoplocephalus . Royal Tyrrell Museum of Palaeontology | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Thyreophora |
Clade: | † Ankylosauria |
Family: | † Ankylosauridae |
Subfamily: | † Ankylosaurinae Brown, 1908 [1] |
Genera [1] | |
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Ankylosaurinae is a subfamily of ankylosaurid dinosaurs, existing from the Early Cretaceous about 105 million years ago until the end of the Late Cretaceous, about 66 mya. Many genera are included in the clade, such as Ankylosaurus , Pinacosaurus , Euoplocephalus , and Saichania .
Ankylosaurinae is formally defined in the PhyloCode as "the largest clade containing Ankylosaurus magniventris , but not Shamosaurus scutatus ". [2] [3] [4] The tribe Ankylosaurini is defined in the PhyloCode as "the largest clade containing Ankylosaurus magniventris , but not Pinacosaurus grangeri and Saichania chulsanensis ". [2]
Diagnostic features of ankylosaurines include the nuchal shelf that obscures the occiput in dorsal view, and the quadrate condyle which is obscured lightly by the quadratojugal boss. [3]
The following cladogram is based on the 50% majority rule phylogenetic analysis of Arbour & Currie (2015): [1]
Thyreophora is a group of armored ornithischian dinosaurs that lived from the Early Jurassic until the end of the Cretaceous.
Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are known to have first appeared in North Africa during the Middle Jurassic, and persisted until the end of the Late Cretaceous. The two main families of ankylosaurians, Nodosauridae and Ankylosauridae are primarily known from the Northern Hemisphere, but the more basal Parankylosauria are known from southern Gondwana during the Cretaceous.
Ankylosaurus is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the non-avian dinosaurs. It was named by Barnum Brown in 1908; it is monotypic, containing only A. magniventris. The generic name means "fused" or "bent lizard", and the specific name means "great belly". A handful of specimens have been excavated to date, but a complete skeleton has not been discovered. Though other members of Ankylosauria are represented by more extensive fossil material, Ankylosaurus is often considered the archetypal member of its group, despite having some unusual features.
Pinacosaurus is a genus of ankylosaurid thyreophoran dinosaur that lived in Asia during the Late Cretaceous, mainly in Mongolia and China.
Saichania is a genus of herbivorous ankylosaurid dinosaur from the Late Cretaceous period of Mongolia and China.
Ankylosauridae is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Paleogene extinction event. These animals were mainly herbivorous and were obligate quadrupeds, with leaf-shaped teeth and robust, scute-covered bodies. Ankylosaurids possess a distinctly domed and short snout, wedge-shaped osteoderms on their skull, scutes along their torso, and a tail club.
Nodosauridae is a family of ankylosaurian dinosaurs known from the Late Jurassic to the Late Cretaceous periods in what is now Asia, Europe, North America, and possibly South America. While traditionally regarded as a monophyletic clade as the sister taxon to the Ankylosauridae, some analyses recover it as a paraphyletic grade leading to the ankylosaurids.
Anodontosaurus is an extinct genus of ankylosaurid dinosaurs within the subfamily Ankylosaurinae. It is known from the entire span of the Late Cretaceous Horseshoe Canyon Formation of southern Alberta, Canada, and is also known from the Dinosaur Park Formation. It contains two species, A. lambei and A. inceptus.
Dyoplosaurus is a monospecific genus of ankylosaurid dinosaur from Alberta that lived during the Late Cretaceous in what is now the Dinosaur Park Formation. Dyoplosaurus represents a close relative of Scolosaurus and Anodontosaurus, two ankylosaurids known from the Horseshoe Canyon and Dinosaur Park Formation.
Tianzhenosaurus is a monospecific genus of ankylosaurid dinosaur from the Shanxi Province that lived during the Late Cretaceous in what is now the Huiquanpu Formation. Tianzhenosaurus may represent a junior synonym of Saichania, an ankylosaurine known from the Barun Goyot and Nemegt Formation.
Gobisaurus is an extinct genus of herbivorous basal ankylosaurid ankylosaur from the Lower Cretaceous of China. The genus is monotypic, containing only the species Gobisaurus domoculus.
Neornithischia is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the inside of their lower teeth. The teeth wore unevenly with chewing and developed sharp ridges that allowed neornithischians to break down tougher plant food than other dinosaurs.
Tarchia is a genus of herbivorous ankylosaurid dinosaur from the late Cretaceous of Mongolia.
Nodocephalosaurus is a monospecific genus of ankylosaurid dinosaur from New Mexico that lived during the Late Cretaceous in what is now the De-na-zin member of the Kirtland Formation. The type and only species, Nodocephalosaurus kirtlandensis, is known only from a partial skull. It was named in 1999 by Robert M. Sullivan. Nodocephalosaurus has an estimated length of 4.5 metres and weight of 1.5 tonnes. It is closely related and shares similar cranial anatomy to Akainacephalus.
Polacanthinae is a subfamily of ankylosaurs, most often nodosaurids, from the Late Jurassic through Early Cretaceous of Europe and potentially North America and Asia. The group is defined as the largest clade closer to Polacanthus foxii than Nodosaurus textilis or Ankylosaurus magniventris, as long as that group nests within either Nodosauridae or Ankylosauridae. If Polacanthus, and by extent Polacanthinae, falls outside either family-level clade, then the -inae suffix would be inappropriate, and the proper name for the group would be the informally defined Polacanthidae.
Minotaurasaurus is a monospecific genus of ankylosaurid dinosaur that lived in Mongolia during the Late Cretaceous in what is now the Djadochta Formation. The type and only species, Minotaurasaurus ramachandrani, is known from two skulls, a cervical vertebra and a cervical half ring. It was named and described in 2009 by Clifford Miles and Clark Miles. The first fossils of Minotaurasaurus were illegally exported out of Mongolia.It has been suggested to be a synonym of Tarchia but more recent publications consider it as a distinct genus.
Ahshislepelta is a monospecific genus of ankylosaur dinosaur from New Mexico that lived during the Late Cretaceous in what is now the Hunter Wash Member of the Kirtland Formation. The type and only species, Ahshislepelta minor, is known only from an incomplete postcranial skeleton of a small subadult or adult individual. It was named in 2011 by Michael Burns and Robert M. Sullivan. Based on the size of the humerus, Ahshislepelta is larger than Pinacosaurus mephistocephalus but smaller than Talarurus and Pinacosaurus grangeri.
Crichtonpelta is a genus of extinct herbivorous ankylosaurid dinosaur from the Late Cretaceous (Cenomanian) of China.
Rhabdodontomorpha is a clade of basal iguanodont dinosaurs. This group was named in 2016 in the context of the description, based on Spanish findings of an early member of the Rhabdodontidae. A cladistic analysis was conducted in which it was found that Muttaburrasaurus was the sister species of the Rhabdodontidae sensu Weishampel. Therefore, Paul-Emile Dieudonné, Thierry Tortosa, Fidel Torcida Fernández-Baldor, José Ignacio Canudo and Ignacio Díaz-Martínez defined Rhabdodontomorpha as a nodal clade: the group consisting of the last common ancestor of Rhabdodon priscus Matheron, 1869 and Muttaburrasaurus langdoni Bartholomai and Molnar, 1981; and all its descendants. Within the clade Zalmoxes and Mochlodon are also included. In 2021, Daniel Madzia redefined Rhabdodontomorpha in the PhyloCode as "the largest clade containing Rhabdodon priscus, but not Iguanodon bernissartensis and Hypsilophodon foxii". The clade is characterized by the following synapomorphies:
Parankylosauria is a group of basal ankylosaurian dinosaurs known from the Cretaceous of South America, Antarctica, and Australia. It is thought the group split from other ankylosaurs during the mid-Jurassic period, despite this being unpreserved in the fossil record.