Crichtonpelta Temporal range: Late Cretaceous, ~ | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Thyreophora |
Clade: | † Ankylosauria |
Family: | † Ankylosauridae |
Subfamily: | † Ankylosaurinae |
Genus: | † Crichtonpelta Arbour & Currie, 2015 |
Species: | †C. benxiensis |
Binomial name | |
†Crichtonpelta benxiensis (Lü et al., 2007) | |
Synonyms | |
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Crichtonpelta is a genus of extinct herbivorous ankylosaurid dinosaur from the Late Cretaceous (Cenomanian) of China. [1]
In 2007, Lü Junchang, Ji Qiang, Gao Yubo and Li Zhixin named and described a second species of Crichtonsaurus : Crichtonsaurus benxiensis. The specific name refers to the Benxi Geological Museum. [2]
The holotype, BXGMV0012, is a skull found near Beipiao in a layer of the Sunjiawan Formation probably dating from the early Cenomanian (~99–95 Ma). Specimen BXGMV0012-1, a skeleton lacking the skull, discovered in the same quarry as the holotype, was referred to the species. [2] Furthermore, a skeleton with skull, displayed by the Sihetun Fossil Museum as a Crichtonsaurus bohlini specimen, was in 2014 referred to Crichtonpelta. [3] In 2017, a fourth specimen was described, from the same quarry as the holotype, G20090034, consisting of a skull lacking the front snout. [4]
In 2014, Victoria Arbour concluded that Crichtonsaurus were a nomen dubium . Therefore, she named a separate genus for its second species: Crichtonpelta. [3] The generic name combines a reference to Michael Crichton, the author of Jurassic Park , with a Greek πέλτη, peltè, "small shield". At the time this was an invalid nomen ex dissertatione. However, in 2015, Crichtonpelta was validly named by Arbour and Philip John Currie. The type species is Crichtonsaurus benxiensis; the combinatio nova is Crichtonpelta benxiensis. [1] There was a possibility that, though Crichtonsaurus bohlini was a nomen dubium, its fossil material in fact belonged to Crichtonpelta. Arbour however, noted clear differences in the scapula and humerus between BXGMV0012-1 and LPM 101, a specimen previously referred to Crichtonsaurus bohlini: the scapula of the former has a tab-like acromion and its humerus a much longer deltopectoral crest. [3]
Arbour established several distinguishing traits. One of these was an autapomorphy, unique derived trait: the apex of the (quadrato)jugal, or cheek, horn, is pointing upwards. Also a unique combination of in themselves not unique traits is present. The upper snout armour forms an amorphous mass, not clearly separated into distinctive tiles. The jugal bone is deeper than that of Pinacosaurus . The skull roof is not notched at the lacrimal bone as in Pinacosaurus grangeri. The squamosal horns are shorter than those of Pinacosaurus mephistocephalus. However, these horns are longer and more pointed than those of Gobisaurus or Shamosaurus . The point of the cheek horn is located on the rear edge. The transverse crest on the top of the rear skull has two points. [1]
The holotype of Crichtonpelta is somewhat larger than Crichtonsaurus, itself about three to four metres long. It is uncertain whether Crichtonpelta already possessed a tail club. [3]
Crichtonpelta was, within the Ankylosauridae, placed in the Ankylosaurinae, in a basal position. [1] If correct, this makes it the oldest known ankylosaurine. [3]
Ankylosaurus is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the non-avian dinosaurs. It was named by Barnum Brown in 1908; it is monotypic, containing only A. magniventris. The generic name means "fused" or "bent lizard", and the specific name means "great belly". A handful of specimens have been excavated to date, but a complete skeleton has not been discovered. Though other members of Ankylosauria are represented by more extensive fossil material, Ankylosaurus is often considered the archetypal member of its group, despite having some unusual features.
Crichtonsaurus is a genus of herbivorous ankylosaurid dinosaur that lived during the Late Cretaceous in what is now China. It was named after Michael Crichton, the author of the dinosaur novel Jurassic Park. A sister taxon was discovered, C. benxiensis, which is now identified as a separate genus.
Pinacosaurus is a genus of ankylosaurid thyreophoran dinosaur that lived in Asia during the Late Cretaceous, mainly in Mongolia and China.
Saichania is a genus of herbivorous ankylosaurid dinosaur from the Late Cretaceous period of Mongolia and China.
Maleevus is an extinct genus of herbivorous ankylosaurid dinosaur from the late Cretaceous, around 90 million years ago, of Mongolia.
Cedarpelta is an extinct genus of basal ankylosaurid dinosaur from Utah that lived during the Late Cretaceous period in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, Cedarpelta bilbeyhallorum, is known from multiple specimens including partial skulls and postcranial material. It was named in 2001 by Kenneth Carpenter, James Kirkland, Don Burge, and John Bird. Cedarpelta has an estimated length of 7 metres and weight of 5 tonnes (11,023 lbs). The skull of Cedarpelta lacks extensive cranial ornamentation and is one of the only known ankylosaurs with individual skull bones that are not completely fused together.
Bissektipelta is a genus of ankylosaurine thyreophoran dinosaurs that lived in Asia during the Late Cretaceous in what is now the Bissekty Formation of Uzbekistan. Bissektipelta is a monospecific genus, containing only the type species B. archibaldi.
Tianzhenosaurus is a genus of ankylosaurid dinosaurs known from the Late Cretaceous Huiquanpu Formation of Shanxi Province, China. The genus contains two species, T. youngi and T. chengi. Some researchers have suggested that Tianzhenosaurus may represent a junior synonym of Saichania, an ankylosaurine from the Barun Goyot and Nemegt formations.
Gobisaurus is an extinct genus of herbivorous basal ankylosaurid ankylosaur from the Lower Cretaceous of China. The genus is monotypic, containing only the species Gobisaurus domoculus.
Tarchia is a genus of herbivorous ankylosaurid dinosaur from the late Cretaceous of Mongolia.
Talarurus is a genus of ankylosaurid dinosaur that lived in Asia during the Late Cretaceous period, about 96 million to 89 million years ago. The first remains of Talarurus were discovered in 1948 and later described by the Russian paleontologist Evgeny Maleev with the type species T. plicatospineus. It is known from multiple yet sparse specimens, making it one of the most well known ankylosaurines, along with Pinacosaurus. Elements from the specimens consists of various bones from the body; five skulls have been discovered and assigned to the genus, although the first two were very fragmented.
Heishansaurus, meaning "Heishan lizard" after the area in China where it was discovered, is the name given to a dubious genus of herbivorous ornithischian dinosaur, probably belonging to the Ankylosauridae.
Shanxia is a monospecific genus of ankylosaurid dinosaur from the Shanxi Province that lived during the Late Cretaceous in what is now the Huiquanpu Formation. Shanxia may possibly represent a junior synonym of Tianzhenosaurus, an ankylosaurine also known from the Huiquanpu Formation of China.
Zhongyuansaurus is a monospecific genus of ankylosaurid dinosaur from Henan that lived during the Early Cretaceous in what is now the Haoling Formation. Zhongyuansaurus is possibly a junior synonym of Gobisaurus, a basal ankylosaurid from the Ulansuhai Formation of Inner Mongolia.
Zhejiangosaurus is an extinct genus of ankylosaurian dinosaur from the Upper Cretaceous of Zhejiang, eastern China. It was first named by a group of Chinese authors Lü Junchang, Jin Xingsheng, Sheng Yiming and Li Yihong in 2007 and the type species is Zhejiangosaurus lishuiensis. It has no diagnostic features, and thus is a nomen dubium.
Ahshislepelta is a monospecific genus of ankylosaur dinosaur from New Mexico that lived during the Late Cretaceous in what is now the Hunter Wash Member of the Kirtland Formation. The type and only species, Ahshislepelta minor, is known only from an incomplete postcranial skeleton of a small subadult or adult individual. It was named in 2011 by Michael Burns and Robert M. Sullivan. Based on the size of the humerus, Ahshislepelta is larger than Pinacosaurus mephistocephalus but smaller than Talarurus and Pinacosaurus grangeri.
Oohkotokia is a genus of ankylosaurid dinosaur within the subfamily Ankylosaurinae. It is known from the upper levels of the Two Medicine Formation of Montana, United States. The discovery of Oohkotokia supports that Ankylosaurine dinosaurs existed and flourished continuously in Montana and/or Alberta throughout the late Campanian and early Maastrichtian stages in the Late Cretaceous period. It was a large, heavily built, quadrupedal, herbivore, that could grow up to 5 metres (16 ft) long and weigh up to 2 metric tons.
Ziapelta is an extinct genus of ankylosaurid. Its fossils have been found in the Hunter Wash and De-na-zin members of the Kirtland Formation of Upper Cretaceous (Campanian) New Mexico. It was named in 2014, in a research paper led by ankylosaur researcher Victoria Arbour. There is a single species in the genus, Ziapelta sanjuanensis. The genus is named after the Zia sun symbol, a stylized sun with four groups of rays, having religious significance to the Zia people of New Mexico, and the iconic symbol on the state flag of New Mexico, and pelta (Latin), a small shield, in reference to the osteoderms found on all ankylosaurids. The specific name is in reference to San Juan County and the San Juan basin, where the fossils were found. Multiple specimens have been described to date, though the fossils are mostly from the front part of the animal. Its closest relative appears to be either Scolosaurus or Nodocephalosaurus, depending on what cladistic model is used.
Zaraapelta is an extinct genus of herbivorous ankylosaurid thyreophoran dinosaur from the Late Cretaceous of Mongolia. The type species is Zaraapelta nomadis, named and described by Arbour et al in 2014. Zaraapelta is known from a single skull from the Barun Goyot Formation. It was found to be closest to Tarchia in the phylogenetic analysis within its description.
Jinyunpelta is a genus of herbivorous ankylosaurine thyreophoran dinosaur from the Cretaceous Liangtoutang Formation of Jinyun County, Zhejiang, China; it has one species, the type species J. sinensis. This species is the basalmost ankylosaur known to have had a proper tail club.