Anoplosaurus Temporal range: late Albian, | |
---|---|
Holotype vertebrae of A. curtonotus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Thyreophora |
Suborder: | † Ankylosauria |
Clade: | † Euankylosauria |
Family: | † Nodosauridae |
Genus: | † Anoplosaurus |
Type species | |
†Anoplosaurus curtonotus |
Anoplosaurus (meaning "unarmored or unarmed lizard") is an extinct genus of herbivorous nodosaurid dinosaur, from the late Albian-age Lower Cretaceous Cambridge Greensand of Cambridgeshire, England. It has in the past been classified with either the armored dinosaurs or the ornithopods, but current thought has been in agreement with the "armored dinosaur" interpretation, placing it in the Ankylosauria.
Harry Govier Seeley named this genus in 1879 for a disarticulated partial postcranial skeleton that had been uncovered no later than 1878 at Reach, Cambridgeshire, composed of a left dentary fragment, numerous vertebrae from the neck, back, and sacrum, parts of the pectoral girdle, humerus fragments, part of the left femur, left tibia, foot bones, ribs, and other fragments. He regarded it as possibly juvenile, due to its small size, [1] with a length of about five feet. The type species is Anoplosaurus curtonotus. The generic name, derived from the Greek hoplo~, a word element used in combinations, with the meaning of "armed", refers to the fact no armour plates had been discovered. The specific name is derived from Latin curtus, "short", and Greek νῶτον, noton, "back".
A second species, Anoplosaurus major, "the larger one", was named by Seeley in 1879 for a neck vertebra and three partial caudal vertebrae he removed from the material previously referred to Acanthopholis stereocercus, from the same formation as the type species. [1] This species now appears to be chimeric, the neck vertebra coming from an ankylosaur, the caudals from an indeterminate iguanodont. [2] [3] [4]
Although Seeley assigned Anoplosaurus to a general Dinosauria, he understood its possible affinities with Scelidosaurus or Polacanthus , as shown by the genus name, and other workers began to see it as an armored dinosaur. [5] [6] In 1902, Baron Franz Nopcsa referred both species to Acanthopholis , creating a Acanthopholis curtonotus and a Acanthopholis major. [7] In 1923 Nopcsa suggested that, while some of the remains belonged to Acanthopholis, other remains, which he removed from that genus, belonged to a camptosaurid. [8] This suggestion led to considerable confusion, with some authors beginning to classify Anoplosaurus under the Camptosauridae, [9] a practice that was continued over several decades (with modifications as iguanodontian taxonomy changed over the years). [10]
In 1964, Oskar Kuhn renamed Syngonosaurus macrocercus Seeley 1879 into Anoplosaurus macrocercus. In 1969, Rodney Steel renamed Eucercosaurus tanyspondylus Seeley 1879 into Anoplosaurus tanyspondylus. Both Syngonosaurus and Eucercosaurus are today seen as nomina dubia and these last two Anoplosaurus species are hereby equally invalid. [2]
In 1998, Xabier Pereda-Suberbiola and Paul Barrett reexamined the material of Anoplosaurus curtonotus. They wrote that it all belonged to a "primitive" or basal member of the Nodosauridae, the lack of armor possibly due to the young age of the animal at death. The basal position would be indicated by the long tooth row and the low sacral vertebrae count. Seeley had never indicated a holotype among the syntype series. Pereda-Superbiola & Barrett therefore selected specimen SMC B55731, a right scapula piece, as the lectotype. Its ankylosaurian affinities would be proven by a high acromion process. The other nodosaurid fossils found at Reach, specimens SMC B55670 - 55742, were assigned as paralectotypes. Pereda-Superbiola & Barrett considered it possible that the discovery had in fact not been made in the Cambridge Green Sand but in the, also Albian, Upper Gault Clay, because the skeletal elements seemed to have belonged to a single individual which might preclude a provenance from the very reworked marine Green Sand deposits. Anoplosaurus curtonotus was by them considered a possibly valid taxon. [2] Reviews since then have followed this interpretation of the genus as an armored dinosaur belonging to the Ankylosauria. [11] [3]
As a possible nodosaurid, Anoplosaurus would have been a quadrupedal, low-slung herbivore, with armour on its body for protection. [3]
Thyreophora is a group of armored ornithischian dinosaurs that lived from the Early Jurassic until the end of the Cretaceous.
Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are known to have first appeared in the Middle Jurassic, and persisted until the end of the Cretaceous Period. The two main families of Ankylosaurs, Nodosauridae and Ankylosauridae are primarily known from the Northern Hemisphere, but the more basal Parankylosauria are known from southern Gondwana during the Cretaceous.
Polacanthus, deriving its name from the Ancient Greek polys-/πολύς- "many" and akantha/ἄκανθα "thorn" or "prickle", is an early armoured, spiked, plant-eating ankylosaurian dinosaur from the early Cretaceous period of England.
Acanthopholis is a genus of ankylosaurian dinosaur in the family Nodosauridae that lived during the Late Cretaceous Period of England. A single species, A. horrida, exists.
Struthiosaurus is a genus of nodosaurid dinosaurs, from the Late Cretaceous period (Santonian-Maastrichtian) of Austria, Romania, France and Hungary in Europe. It was a small dinosaur, measuring 2–3 m (6.6–9.8 ft) in length and weighing 300–400 kg (660–880 lb).
Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia.
Stegopelta is a genus of struthiosaurin nodosaurid dinosaur. It is based on a partial skeleton from the latest Albian-earliest Cenomanian-age Lower and Upper Cretaceous Belle Fourche Member of the Frontier Formation of Fremont County, Wyoming, USA.
Syngonosaurus is an extinct genus of ornithopod dinosaur from the Early Cretaceous. It was an iguanodontian discovered in England and was first described in 1879. The type species, S. macrocercus, was described by British paleontologist Harry Seeley in 1879 and it was later synonymised with Acanthopholis, but the genus was reinstated in a 2020 study, when Syngonosaurus and Eucercosaurus were reinterpreted as basal iguanodontians.
Texasetes is a genus of ankylosaurian dinosaurs from the late Lower Cretaceous of North America. This poorly known genus has been recovered from the Paw Paw Formation near Haslet, Tarrant County, Texas, which has also produced the nodosaurid ankylosaur Pawpawsaurus.
Cedarpelta is an extinct genus of basal ankylosaurid dinosaur from Utah that lived during the Late Cretaceous period in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, Cedarpelta bilbeyhallorum, is known from multiple specimens including partial skulls and postcranial material. It was named in 2001 by Kenneth Carpenter, James Kirkland, Don Burge, and John Bird. Cedarpelta has an estimated length of 7 metres and weight of 5 tonnes (11,023 lbs). The skull of Cedarpelta lacks extensive cranial ornamentation and is one of the only known ankylosaurs with individual skull bones that are not completely fused together.
Hoplitosaurus was a genus of armored dinosaur related to Polacanthus. It was named from a partial skeleton found in the ?Barremian-age Lower Cretaceous Lakota Formation of Custer County, South Dakota. It is an obscure genus which has been subject to some misinterpretation of its damaged remains. Although there was a push to synonymize it with Polacanthus in the late 1980s-early 1990s, Hoplitosaurus has been accepted as a valid albeit poorly known genus in more recent reviews.
Denversaurus is a genus of panoplosaurin nodosaurid dinosaur from the late Maastrichtian of Late Cretaceous Western North America. Although at one point treated as a junior synonym of Edmontonia by some taxonomists, current research indicates that it is its own distinct nodosaurid genus.
Silvisaurus, from the Latin silva "woodland" and Greek sauros "lizard", is a nodosaurid ankylosaur from the Early to Late Cretaceous period.
Dracopelta is a monospecific genus of ankylosaur dinosaur from Portugal that lived during the Late Jurassic in what is now the Lourinhã Formation. The type and only species is Dracopelta zbyszewskii, which is represented by a partial skeleton including unpublished material.
Eucercosaurus is the name given to a genus of ornithopod dinosaur from the Albian stage of the Early Cretaceous. It was an ornithopod discovered in the Cambridge Greensand of England and is known from 19 centra, 3 sacrals, 4 dorsals and 12 caudals, and a neural arch found near Trumpington, Cambridgeshire. The type species, E. tanyspondylus, was described by British paleontologist Harry Seeley in 1879. It is considered a dubious name, and was once considered an ankylosaur. According to a 2020 study, Eucercosaurus and Syngonosaurus were basal iguanodontians.
Glyptodontopelta is a monospecific genus of nodosaurid dinosaur from New Mexico that lived during the Late Cretaceous in what is now the Naashoibito member of the Ojo Alamo Formation. The type and only species, Glyptodontopelta mimus, is known from numerous specimens that consist of osteoderms, a dentary, supraorbital and bone fragments. It was named in 2000 by Tracy Ford. Edmontonia australis is a junior synonym of Glyptodontopelta.
Priodontognathus was a genus of ankylosaurian dinosaur possibly from the Oxfordian-age Upper Jurassic Lower Calcareous Grit of Yorkshire, England. It is a dubious genus based on a maxilla, and has been erroneously mixed up with iguanodonts and stegosaurs.
The Cambridge Greensand is a geological unit in England whose strata are earliest Cenomanian in age. It lies above the erosive contact between the Gault Formation and the Chalk Group in the vicinity of Cambridgeshire, and technically forms the lowest member bed of the West Melbury Marly Chalk Formation. It is a remanié deposit, containing reworked fossils of late Albian age, including those of dinosaurs and pterosaurs.
This timeline of ankylosaur research is a chronological listing of events in the history of paleontology focused on the ankylosaurs, quadrupedal herbivorous dinosaurs who were protected by a covering bony plates and spikes and sometimes by a clubbed tail. Although formally trained scientists did not begin documenting ankylosaur fossils until the early 19th century, Native Americans had a long history of contact with these remains, which were generally interpreted through a mythological lens. The Delaware people have stories about smoking the bones of ancient monsters in a magic ritual to have wishes granted and ankylosaur fossils are among the local fossils that may have been used like this. The Native Americans of the modern southwestern United States tell stories about an armored monster named Yeitso that may have been influenced by local ankylosaur fossils. Likewise, ankylosaur remains are among the dinosaur bones found along the Red Deer River of Alberta, Canada where the Piegan people believe that the Grandfather of the Buffalo once lived.