Cleftbelly trevally

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Cleftbelly trevally
Cleftbelly trevally 2.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Carangiformes
Family: Carangidae
Subfamily: Caranginae
Genus: Atropus
Oken, 1817
Species:
A. atropos
Binomial name
Atropus atropos
(Bloch & Schneider, 1801)
Atropus atropos distribution.png
Approximate range of the cleftbelly trevally
Synonyms
  • Brama atropos,
    Bloch & Schneider, 1801
  • Caranx atropus,
    (Bloch & Schneider, 1801)
  • Atropus atropus(misspelling)

The cleftbelly trevally (Atropus atropos), also known as the cleftbelly kingfish, Kuweh trevally or thin crevalle, is a species of tropical marine fish of the jack family, Carangidae. The species inhabits coastal waters throughout the Indo-West Pacific region from South Africa in the west to Japan in the east, often found near the water's surface. The cleftbelly trevally is the only member of the genus Atropus and is distinguished by a number of anatomical characteristics, with a deep median groove in the belly giving the species its common name. It is not a large fish, growing to a maximum recorded length of 26.5 cm. Cleftbelly trevally are predatory fish, taking a variety of small crustaceans and fish. The species is of minor importance to fisheries throughout its range.

Contents

Taxonomy and naming

The cleftbelly trevally is the only species in the genus Atropus, which is one of 31 genera in the jack family, Carangidae. The Carangidae are part of the order Carangiformes. [2]

The species was first scientifically described under the name of Brama atropos by German ichthyologists Marcus Elieser Bloch and Johann Gottlob Schneider in the massive 1801 volume of Systema Ichthyologiae iconibus cx illustratum, a document which is the taxonomic authority for many species. [3] Bloch and Schneider placed the species in the genus Brama , within the pomfret family. The generic position of the species was revised twice, once being placed in the jack genus, Caranx , and finally into Atropus. The genus Atropus had been informally created by Georges Cuvier in 1817 as "Les Atropus", and was formally Latinized by Lorenz Oken, thus making him the author of the genus. [4] A. atropos is the type species of the genus by monotypy. The type specimen was taken from the waters of Puducherry, in India.

The fact that the generic and specific names differ by a single letter has also led to the misspelling of the species as Atropus atropus in some publications. [4] The species is most commonly referred to in English as the cleftbelly trevally, with other names including cleftbelly kingfish, Kuweh trevally and thin crevalle. [3] It is also occasionally called the 'blackfin jack', but should not be confused with Hemicaranx zelotes , also known by that name.

Description

Male cleftbelly trevally Atropus atropos.png
Male cleftbelly trevally

The cleftbelly trevally has a body shape similar in many ways to a number of the larger jacks, with a strongly compressed body almost ovate in shape. [5] The dorsal profile of the head is straight and steep from the tip of the upper jaw to above the eye where the profile becomes convex. The eye lacks an adipose eyelid, and the upper jaw has a narrow band of small teeth, while the lower jaw has two or three rows of small teeth which narrow to one row on the sides of the jaw. [5] The ventral profile is concave upwards, with a major distinguishing feature of the species being a deep median groove along the belly, into which the large pelvic fins fold, also encompassing the first two anal fin spines and anus. The entire chest from underneath the pectoral fins to the base of the pelvic fins is scaleless. [4] There are two separate dorsal fins, the first with eight spines and the second with a single spine followed by 19 to 22 soft rays. [6] The anal fin has two detached spines before the major part of the anal fin, which consists of one spine and 17 or 18 soft rays. The pectoral and pelvic fins are both quite long, with the latter extending almost to the anal fin origin. The lateral line has a moderately strong arch anteriorally which intersects the straight section underneath the fifth to seventh dorsal rays. The straight part of the lateral line has 31–37 scutes. [6]

A cleftbelly trevally at an Indian market Cleftbelly trevally.jpg
A cleftbelly trevally at an Indian market

There are 24 vertebrae and 29 to 34 total gill rakers. [5] The species is known to reach 26.5 cm in length, although often encountered below 20 cm. [3]

The cleftbelly trevally is a bluish green on the upper body which fades to silver on the underside of the fish. The membranes of the large pelvic fin are a distinctive black, while the remaining fins are a pale white colour. Juveniles have indistinct dark bands across their bodies, with a black spot becoming more prominent in adults. [6]

The species show sexual dimorphism in their dorsal fins, with mature males having between 6 and 12 elongated soft rays, strung out into filaments of varying lengths. The anal fin is also has five similar extensions of the soft rays. Females do not show this elongation. [7]

Distribution and habitat

The cleftbelly trevally inhabits the tropical and subtropical waters of the Indo-West Pacific region. Its range extends from South Africa [8] along the east African coast [9] including the Persian Gulf, east to India, Southeast Asia, the Philippines, Taiwan and southern Japan. [5] Some sources state the Persian Gulf as the westernmost point of the species range, but specimens sent to Henry Fowler from South and eastern Africa in the 1930s show this to be incorrect. [8] [9]

The species inhabit shallow coastal waters, where they swim close to the surface. There has been a report of the species living in areas near estuaries which empty large flows of fresh water into these areas, indicating the species may be able to live in estuarine environments. [10]

Biology and fishery

The cleftbelly trevally is a predatory species, preying on small crustaceans including shrimps, copepods and decapods, as well as small fish. [11] Reproduction in the species has not been extensively studied, although early growth has been described in a 1984 article. The study was located in India, and showed the fish grows 12.4 cm in its first year of life, growing slower in its second and third years, adding 7.35 cm and 3.5 cm to its total length, respectively, in these years. [12] The species reaches sexual maturity at 21 cm, in the second or third year of its life. [13]

The species is of minor importance to fisheries throughout its range, taken mainly by trawls, traps and by various hook-and-line methods. [5]

Related Research Articles

<span class="mw-page-title-main">Crevalle jack</span> Species of fish

The crevalle jack, also known as the common jack, black-tailed trevally, couvalli jack, black cavalli, jack crevale, or yellow cavalli is a common species of large marine fish classified within the jack family, Carangidae. The crevalle jack is distributed across the tropical and temperate waters of the Atlantic Ocean, ranging from Nova Scotia, Canada to Uruguay in the western Atlantic and Portugal to Angola in the eastern Atlantic, including the Mediterranean Sea. It is distinguishable from similar species by its deep body, fin colouration and a host of more detailed anatomical features, including fin ray and lateral line scale counts. It is one of the largest fish in the genus Caranx, growing to a maximum known length of 124 cm and a weight of 32 kg, although is rare at lengths greater than 60 cm. The crevalle jack inhabits both inshore and offshore waters to depths of around 350 m, predominantly over reefs, bays, lagoons and occasionally estuaries. Young fish dispersed north by currents in the eastern Atlantic are known to migrate back to more tropical waters before the onset of winter; however, if the fish fail to migrate, mass mortalities occur as the temperature falls below the species' tolerance.

<span class="mw-page-title-main">Bigeye trevally</span> Species of fish

The bigeye trevally, also known as the bigeye jack, great trevally, six-banded trevally and dusky jack, is a species of widespread large marine fish classified in the jack family Carangidae. The bigeye trevally is distributed throughout the tropical waters of the Indian and Pacific Oceans, ranging from South Africa in the west to California and Ecuador in the east, including Australia to the south and Japan in the north. The bigeye trevally is best distinguished by its colouration, having a dark second dorsal fin with a white tip on the lobe, and also possessing a small dark spot on the operculum. Other more detailed anatomical features also set the species apart from other members of Caranx. The species is known to grow to a length of 120 cm and 18 kg.

<span class="mw-page-title-main">African pompano</span> Species of fish

The African pompano, also known as the pennant-fish or threadfin trevally, is a widely distributed species of tropical marine fish in the jack family, Carangidae. The species is found in tropical waters worldwide, with adults often inhabiting coastlines, while juveniles are usually pelagic, floating with ocean currents. The adult African pompano is similar in appearance to the other members of the genus Alectis, with the concave shape of the head near the eyes; the clearest distinguishing feature. The juveniles are similar to other members of Alectis, having long, filamentous dorsal and anal fin tips which are thought to discourage predators. The species lives in depths less than 100 m, consuming a range of crustaceans and small fishes. The species is of minor economic importance, often taken amongst other tropical midwater fishes by hook and line, while juveniles are occasionally caught in beach seines. African pompano are also highly rated game fish, often considered one of the strongest of the jacks in larger sizes.

<span class="mw-page-title-main">Malabar trevally</span> Species of fish

The Malabar trevally, also known as the Malabar jack, Malabar kingfish or nakedshield kingfish, is a species of large inshore marine fish of the jack family, Carangidae. It is distributed throughout the Indian and west Pacific Oceans from South Africa in the west to Japan and Australia in the east, inhabiting reefs and sandy bays on the continental shelf. The Malabar trevally is similar to many of the other species in the genus Carangoides, with the number of gill rakers and the grey-brown colour of the tongue being the diagnostic features. The Malabar trevally is a predator, taking a variety of small fish, cephalopods and crustaceans. The species is of minor economic importance throughout its range, caught by a variety of net and handline methods.

<span class="mw-page-title-main">Herring scad</span> Species of fish

The herring scad, is a common species of tropical marine fish in the jack family Carangidae. The species inhabits the surface waters of coastal regions throughout the Indo-West Pacific region, feeding on a variety of crustaceans and small fishes. It is the largest fish of the scad genus Alepes, growing to a recorded length of 56 cm. The herring scad is identified among the genus Alepes by its more numerous and smaller scutes and the number of gill rakers on the first arch. It is of minor importance to fisheries throughout its range.

<span class="mw-page-title-main">Longfin trevally</span> Species of fish

The longfin trevally, also known as the longfin kingfish, longfin cavalla or armed trevally, is a species of inshore marine fish in the jack family, Carangidae. The species is common in tropical to subtropical waters of the Indo-Pacific, ranging from South Africa in the west to Japan in the east, typically inhabiting inshore reefs and bays. The species is easily distinguished by its elongate dorsal and anal fin lobes and filamentous dorsal rays, as well as its scaleless breast. Longfin trevally are pelagic predators, taking a variety of small fish, cephalopods and crustaceans, and reach sexual maturity at around 21 cm. The species has a maximum known length of 57 cm and weight of 3.5 kg. The longfin trevally has a very complex taxonomic history which is closely intertwined with another currently valid species, Carangoides ciliarius, which may yet prove to be synonymous. Longfin trevally are of minor importance to fisheries throughout their range and are considered good table fish, and are occasionally taken by anglers.

<span class="mw-page-title-main">Longnose trevally</span> Species of fish

The longnose trevally, also known as the tea-leaf trevally, club-nosed trevally, grunting trevally or dusky trevally, is a species of inshore marine fish in the jack family, Carangidae. The species is distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans from South Africa to New Zealand and Japan, inhabiting coastal waters, especially reefs, to a depth of 90 m. The longnose trevally is distinguished from similar species by a combination of a scaleless breast and the number of gill rakers and fin rays. It is a moderately large fish, growing to a maximum known length of 72 cm and 4.35 kg. The longnose trevally is a predatory fish, consuming small fish, crustaceans and molluscs. The species is of minor commercial importance throughout its range, and is considered to be a good table fish.

<span class="mw-page-title-main">Shadow trevally</span> Species of fish

The shadow trevally, also known as the shadow kingfish, twothread trevally or Aldabra trevally, is a species of inshore marine fish in the jack family Carangidae. The species is patchily distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans, from South Africa in the west to Japan and Samoa in the east, reaching as far south as Indonesia and New Caledonia. It is most easily distinguished from similar species by as series of dark rectangular blotches under the second dorsal fin, giving a 'shadowed' appearance, from which its common name is derived. The shadow trevally is a reasonably large fish, growing to 85 cm in length and at least 2.6 kg in weight. It inhabits shallow coastal waters, including reefs, bays, and estuaries, where it takes small fish and benthic crustaceans as prey. Nothing is known of the species' ecology and reproductive biology. It is of little importance to fisheries, and is occasionally taken by bottom trawls and other artisanal fishing gear.

<i>Caranx</i> Genus of fishes

Caranx is a genus of tropical to subtropical marine fishes in the jack family Carangidae, commonly known as jacks, trevallies and kingfishes. They are moderate- to large-sized, deep-bodied fishes which are distinguished from other carangid genera by specific gill raker, fin ray and dentition characteristics. The genus is represented in the Pacific, Indian and Atlantic Oceans, inhabiting both inshore and offshore regions, ranging from estuaries and bays to deep reefs and offshore islands. All species are powerful predators, taking a variety of fish, crustaceans and cephalopods, while they in turn are prey to larger pelagic fishes and sharks. A number of fish in the genus have a reputation as powerful gamefish and are highly sought by anglers. They often make up high amounts of the catch in various fisheries, but are generally considered poor to fair table fishes.

<span class="mw-page-title-main">Whitefin trevally</span> Species of fish

The whitefin trevally, also known as the horse trevally, is a species of deep water offshore fish in the jack family Carangidae. The species inhabits the tropical to temperate waters of the Indo-Pacific and central Pacific, ranging from South Africa in the west to Hawaii in the east. The whitefin trevally is a moderate-sized fish, growing to 37 cm, and is distinguished by a number of morphological traits, including fin size, gill raker count, and colour. It inhabits the continental shelf and slope at depths to 200 m over sand and mud substrates, where it preys on fish, crustaceans, and cephalopods. Studies in Japan indicate a length at sexual maturity of 17.4 cm on average, with spawning occurring between May and October, with each individual spawning multiple times. Whitefin trevallies are of high importance to fisheries in Japan, where they are taken by trawlers, although the catch numbers have halved since the 1980s. It is of minor importance elsewhere throughout its range, but is considered a good table fish.

<span class="mw-page-title-main">Blue trevally</span> Species of fish

The blue trevally, also known as the banded trevally, barred trevally, Ferdau's trevally or Forskaal's jackfish, is a common, widespread species of pelagic marine fish classified in the jack family, Carangidae. The blue trevally is distributed throughout the tropical and subtropical waters of the Indo-Pacific and central Pacific regions, ranging from South Africa in the west to Hawaii in the east. It is a moderately large fish, growing to a recorded maximum length of 70 cm, with the number of rays in the second dorsal fin and the colouring serving as diagnostic features of the species. The species inhabits waters to depths of 60 m, generally inhabiting reefs, beaches, lagoons, and areas with sandy substrates. It is a predatory fish, taking other fish, prawns, crabs, and molluscs, and very little is known of the species' reproductive biology. The blue trevally is of varying importance to fisheries throughout its range, with some regions having high catches of the fish. It is considered to be a gamefish, and is sought after for its excellent eating qualities.

<span class="mw-page-title-main">Yellowspotted trevally</span> Species of fish

The yellowspotted trevally, also known as the yellowspotted kingfish, goldspotted trevally, tarrum, or yellowspot, is a widespread species of large inshore marine fish in the jack family Carangidae. The yellowspotted trevally inhabits the tropical and subtropical waters of the western Indo-Pacific region, from South Africa in the west to Japan and Australia in the east. The species is known to grow to a maximum length of at least 1.2 m, and is distinguished by gill raker and fin morphology, as well as the distinctive golden spots which give the fish its name. The yellowspotted trevally generally prefers inshore rocky and coral reefs, but is occasionally found over deep offshore sand banks to a depth of 100 m. It is a predatory fish, taking fish, cephalopods, and crustaceans, and shows diet partitioning with other trevallies in studies conducted in Australian waters. Reproduction is poorly studied, although observational evidence suggests spawning occurs in aggregations, probably during summer in South Africa. It is generally of minor importance to commercial fisheries throughout its range, but is considered an excellent sportfish by anglers and spearfishermen, and a good table fish.

<span class="mw-page-title-main">Bumpnose trevally</span> Species of fish

The bumpnose trevally, also known as the bumpnose kingfish or onion kingfish, is a species of relatively small inshore marine fish classified in the jack family Carangidae. The bumpnose trevally is fairly common in the tropical and subtropical waters of the Indo-west Pacific region, ranging from South Africa in the west to Japan and Samoa in the east. It is a small species by carangid standards, reaching a maximum known length of 32 cm, and can be distinguished from the similarly shaped Carangoides armatus by a distinct 'bump' on the snout, which gives the fish its common name. The species inhabits coastal waters, often living along bays and beaches, where it takes shrimp, small crabs, and juvenile fish as prey. The bumpnose trevally is of minor importance to fisheries throughout its range, taken by hook and line, trawls, and seine nets. It is also of minor importance to anglers, taken by baits from beaches and piers, and is considered a modest table fish.

<span class="mw-page-title-main">Duskyshoulder trevally</span> Species of fish

The duskyshoulder trevally or epaulet trevally, is a species of small inshore marine fish in the jack family, Carangidae. It is distributed through the eastern Indian and western Pacific Oceans, ranging from eastern India to northern Australia and Taiwan. It is relatively small by carangid standards, reaching only 27 cm maximum length, and can be distinguished by the large, black blotches on its shoulders. The duskyshoulder trevally is an inshore fish living in waters less than 50 m deep, over sandy substrates in bays and on the continental shelf. It is a predatory fish, taking demersal fishes, crustaceans, and cephalopods, with nothing known of its reproductive habits. It is of little value to fisheries, often taken as bycatch in prawn trawling operations.

<span class="mw-page-title-main">Coachwhip trevally</span> Species of fish

The coachwhip trevally, also known as the oblong trevally or oblique-banded trevally, is a species of inshore marine fish classified in the jack family Carangidae. The coachwhip trevally is distributed through the Indo-west Pacific region, ranging from South Africa in the west to Fiji and Japan in the east. It is a moderately large fish, growing to a known maximum length of 46 cm and can be distinguished from similar species by an array of detailed morphological features including dentition, fin ray counts and scale patterns. The coachwhip trevally inhabits coastal waters throughout its range, known to prefer estuarine waters in a number of localities. Nothing is known of its diet or reproductive biology, and is of little importance to fisheries, occasionally taken as bycatch in trawl and hook and line fisheries.

<span class="mw-page-title-main">Imposter trevally</span> Species of fish

The imposter trevally, also known as the imposter jack or white-tongued trevally, is a species of small coastal marine fish in the jack family Carangidae. The imposter trevally is distributed through the tropical waters of Indian and west Pacific oceans, from the Gulf of Oman in the west to Japan and Australia in the east. The species is quite similar to the Malabar trevally, but can be distinguished by its gill raker count, and is identifiable from other species by having a white to pale grey tongue. It is a relatively small species, growing to a maximum recorded length of 30 cm. The imposter trevally inhabits coastal waters of the continental shelf in depths of up to 140 m, often associating with closely related carangid species. It preys on a range of small fish, crustaceans and cephalopods, but little is known of its reproductive biology. The imposter trevally is of minor importance to fisheries throughout its range, taken by hook and line, bottom trawls and several types of artisanal gear.

<span class="mw-page-title-main">Longfin crevalle jack</span> Species of fish

The longfin crevalle jack is a recently described species of large marine fish classified in the jack family, Carangidae. It inhabits the subtropical waters of the east Atlantic Ocean, ranging along the African coast from Mauritania south at least to Moçamedes in southern Angola, with the species historically present in the Mediterranean Sea. It is an inshore species, known to occasionally penetrate estuaries, possibly to spawn. The species is very similar to the crevalle jack, Caranx hippos, and is separated by its extended dorsal and anal fin lobes as well as more detailed anatomical features including dorsal and anal fin ray counts. The fish is known to reach 127 cm in length and 20.9 kg in weight. The longfin crevalle jack is a predatory fish, taking small fish as its main prey. Due to longstanding confusion between C. fischeri and C. hippos, the importance of each species to African fisheries is poorly understood, with a known combined catch in this region of between 2,233 and 10,054 tonnes per year in 1995–2004. Like its close relatives, the species is considered a powerful gamefish and highly sought after, although their coarse flesh makes for relatively poor quality food.

<span class="mw-page-title-main">Tille trevally</span> Species of fish

The tille trevally, also known as the tille kingfish, is a species of large marine fish classified in the jack family, Carangidae. The tille trevally is distributed through the tropical and subtropical waters of the Indo-Pacific region, ranging from South Africa in the west to Fiji, Japan and Australia in the east. The species is best distinguished by its rounded, strongly convex anterior profile, with other detailed anatomical features also useful. The tille trevally reaches a maximum length of 80 cm and a weight of 7.2 kg. It is predominantly an inshore species, inhabiting coastal reef and lagoon environments, although has been recorded on deep offshore seamounts. It is a predatory fish, taking various species of fish and crustaceans as prey, with little known of its reproductive cycle. It is of minor importance to fisheries throughout its range, taken by hook and line, gill nets and purse seines. The tille trevally is also considered a good game fish, and an excellent table fish. The species acquired its scientific and common names from a local name used by Pondicherry fishermen, koton tille, which Georges Cuvier then used when he named the species in 1833.

<span class="mw-page-title-main">Yellowstripe scad</span> Species of fish

The yellowstripe scad, also known as the yellowstripe trevally, yellow-banded trevally, smooth-tailed trevally, slender-scaled trevally and slender trevally, is a species of small inshore fish in the jack and horse mackerel family Carangidae, and the only member of the genus Selaroides. The yellowstripe trevally is distributed throughout the tropical waters of the Indo-West Pacific region, ranging from the Persian Gulf in the west to Vanuatu and New Caledonia in the east. The species is distinguished by its prominent lateral yellow band, and differs from the scads of the genus Selar in having a smaller eye and different dentition. Although the yellowstripe scad reaches a maximum recorded length of 22 cm, it is normally encountered at sizes less than 15 cm. Phylogenetic studies indicate the yellowstripe scad is closely related to the scads of the genus Selar, although its exact placement in the family Carangidae is less well agreed upon.

<span class="mw-page-title-main">Longrakered trevally</span> Species of fish

The longrakered trevally, also known as the cale cale trevally and heavyjawed kingfish, is a species of marine fish in the jack and horse mackerel family Carangidae. The longrakered trevally is distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans, from Mozambique and Madagascar in the west, to Japan and northern Australia in the east. A large species growing to a recorded length of 1 m, the longrakered trevally is distinguished by is protruding lower jaw, elongated gill rakers and lack of villiform teeth on its tongue. It is an inshore species, restricted to coastal and estuarine regions, where it preys on fishes and crustaceans. Little is known of the species reproductive cycle or growth. The longrakered trevally is of minor importance to fisheries and is often taken as bycatch in finfish and prawn trawls, as well as by recreational fishermen.

References

  1. Smith-Vaniz, W.F.; Borsa, P.; Carpenter, K.E.; Jiddawi, N.; Yahya, S.; Obota, C. (2018). "Atropus atropos". IUCN Red List of Threatened Species . 2018: e.T20256645A46664034. doi: 10.2305/IUCN.UK.2018-2.RLTS.T20256645A46664034.en . Retrieved 20 November 2021.
  2. J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016). Fishes of the World (5th ed.). Wiley. pp. 380–387. ISBN   978-1-118-34233-6. Archived from the original on 2019-04-08. Retrieved 2019-11-23.
  3. 1 2 3 Froese, Rainer; Pauly, Daniel (eds.) (2007). "Atropus atropos" in FishBase . November 2007 version.
  4. 1 2 3 Lin, Pai-Lei; Shao, Kwang-Tsao (18 April 1999). "A Review of the Carangid Fishes (Family Carangidae) From Taiwan with Descriptions of Four New Records". Zoological Studies. 38 (1): 33–68. Archived from the original on 29 February 2012. Retrieved 1 January 2008.
  5. 1 2 3 4 5 Carpenter, Kent E.; Niem, Volker H., eds. (2001). FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Volume 5. Bony fishes part 3 (Menidae to Pomacentridae) (PDF). Rome: FAO. p. 2684. ISBN   92-5-104587-9.
  6. 1 2 3 Randall, John E. (1995). Coastal Fishes of Oman. Honolulu: University of Hawaii Press. p. 183. ISBN   0-8248-1808-3.
  7. Shameen, A.; Dutt, S. (1984). "A note on sexual dimorphism in carangid fishes". Mahasagar. 17 (3): 179–181. ISSN   0542-0938.
  8. 1 2 Fowler, Henry W. (1935). "South African Fishes Received from Mr. H. W. Bell-Marley in 1935". Proceedings of the Academy of Natural Sciences of Philadelphia. 87: 361–408.
  9. 1 2 Fowler, Henry W. (1934). "Fishes Obtained by Mr. H. W. Bell-Marley Chiefly in Natal and Zululand in 1929 to 1932". Proceedings of the Academy of Natural Sciences of Philadelphia. 86: 405–514.
  10. Ansari, Z.A.; Chatterji, A.; Ingole, B. S.; et al. (1995). "Community Structure and Seasonal Variation of an Inshore Demersal Fish Community at Goa, West Coast of India". Estuarine, Coastal and Shelf Science . 41 (5). Elsevier: 593–610. Bibcode:1995ECSS...41..593A. doi:10.1016/0272-7714(95)90029-2.
  11. Masuda, H.; Amaoka, K.; Araga, C.; Uyeno, T.; Yoshino, T. (1984). The fishes of the Japanese Archipelago. Vol. 1. Tokyo, Japan: Tokai University Press. p. 437. ISBN   978-4-486-05054-4.
  12. Kochar, L.S.; S.N. Dwivedi (1988). "A Comparative study of Morphometric Relationships and Age and Growth of Atropus atropus and Carangoides malabaricus Bloch and Schneider off Bombay Coast India". Proceedings of the National Academy of Sciences, India Section B. 58 (4): 515–524. ISSN   0369-8211.
  13. Reuben, S.; Kasim, H.M.; Sivakami, S.; et al. (1992). "Fishery, biology and stock assessment of carangid resources from the Indian seas". Indian Journal of Fisheries. 39 (3–4): 195–234.