Coelacanthus Temporal range: | |
---|---|
Coelacanthus granulatus fossil | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Sarcopterygii |
Class: | Actinistia |
Order: | Coelacanthiformes |
Family: | † Coelacanthidae |
Genus: | † Coelacanthus Agassiz, 1839 |
Species: | †C. granulatus |
Binomial name | |
†Coelacanthus granulatus Agassiz, 1839 | |
Coelacanthus ("hollow spine") is a genus of extinct marine coelacanths known from the late Permian period. It was the first genus of coelacanths described, about a century before the discovery of the extant coelacanth Latimeria . [1] The order Coelacanthiformes is named after it.
The only definitive species in this genus is C. granulatus from the late Permian (Wuchiapingian stage) Kupferschiefer of Germany and equivalent Marl Slate of England. [2] [3]
The genus has long been used to group unrelated species of coelacanths, and several other species that were first referred to Coelacanthus were later reallocated to other genera. [2] Coelacanthus minor was considered by Woodward (1891) as potentially belonging to the Triassic genus Heptanema , [4] while Martin and Wenz (1984) considered Coelacanthus lunzensis a possible synonym of Garnbergia . [5] Coelacanthus madagascariensis from the Early Triassic of Madagascar was reattributed to the genus Rhabdoderma , and Coelacanthus evolutus is a synonym of Whiteia woodwardi . [6]
The following are considered species that likely do not belong to this genus due to either lack of phylogenetic analysis or incomplete remains, but have not yet been reclassified: [2]
Coelacanthus bears a superficial similarity to the living coelacanth Latimeria , though it was smaller, and had a more elongated head. Individuals grew up to 0.7 metres (2.3 ft) in length, had an elongate codavypter or supplementary tail lobe, and had small lobed fins, suggesting that Coelacanthus were open-water predators. The fin rays of the caudal fin are hollow, which gave Coelacanthus its name. The name is an adaptation of the Modern Latin cœlacanthus ("hollow spine"), from the Greek κοῖλ-ος (koilos; "hollow") and ἄκανθ-α (akantha; "spine"). These hollow spines are a typical feature of coelacanths.
Coelacanths are an ancient group of lobe-finned fish (Sarcopterygii) in the class Actinistia. As sarcopterygians, they are more closely related to lungfish and tetrapods than to ray-finned fish.
Semionotiformes is an order of ray-finned fish known from the Middle Triassic (Anisian) to the Late Cretaceous (Maastrichtian). Their closest living relatives are gars (Lepisosteidae), with both groups belonging to the clade Ginglymodi within the Holostei. The group includes both freshwater (Semionotidae) and marine adapted forms. Many members of the family Macrosemiidae, had elongated dorsal fins, often associated with an adjacent area of skin which was free of scales. These fins were likely undulated for use in precision swimming. The body morphology of macrosemiids suggests that they were slow swimmers who were capable of maneuvering around complex topography, such as reef environments.
Chimaeras are cartilaginous fish in the order Chimaeriformes, known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively.
In the geologic timescale, the Wuchiapingian or Wujiapingian is an age or stage of the Permian. It is also the lower or earlier of two subdivisions of the Lopingian Epoch or Series. The Wuchiapingian spans the time between 259.51 and 254.14 million years ago (Ma). It was preceded by the Capitanian and followed by the Changhsingian.
Acanthodes is an extinct genus of acanthodian fish. Species have been found in Europe, North America, and Asia, spanning the Early Carboniferous to the Early Permian, making it one of the youngest known acanthodian genera.
Palaeoniscum is an extinct genus of ray-finned fish from the Permian period (Guadalupian-Lopingian) of England, Germany, Turkey, North America and Greenland, and possibly other regions. The genus was named Palaeoniscum in 1818 by Henri Marie Ducrotay de Blainville, but was later misspelled as Palaeoniscus by Blainville and other authors. Palaeoniscum belongs to the family Palaeoniscidae.
Saurichthys is an extinct genus of predatory ray-finned fish from the Triassic Period. It is the type genus of the family Saurichthyidae, and the most speciose and longest lasting genus in the family. This family also includes the Permian Eosaurichthys (China) and the Jurassic Saurorhynchus from Europe and North America, though it may be more appropriate to treat these as subgenera of Saurichthys, due to the genus Saurichthys otherwise being paraphyletic.
The West Indian Ocean coelacanth is a crossopterygian, one of two extant species of coelacanth, a rare order of vertebrates more closely related to lungfish and tetrapods than to the common ray-finned fishes. The other extant species is the Indonesian coelacanth.
Rhabdoderma is an extinct genus of coelacanth fish in the class Sarcopterygii. It lived in the Carboniferous and Early Triassic (Induan), and its fossils have been found in Europe, Madagascar and North America. The type species was originally described as Coelacanthus elegans. Five species are considered valid in 1981.
Coccoderma is an extinct genus of prehistoric marine coelacanth which lived during the Late Jurassic period. Fossils have been found in Germany and France. It was small in size, about 27.5 cm. They had very long and sharp teeth.
Ticinepomis is an extinct genus of coelacanth lobe-finned fish which lived during the Middle Triassic period in what is now Switzerland. It contains two species, T. peyeri and T. ducanensis.
Mawsonia is an extinct genus of prehistoric coelacanth fish. It is amongst the largest of all coelacanths, with one quadrate specimen possibly belonging to an individual measuring 5.3 metres in length. It lived in freshwater and brackish environments from the late Jurassic to the mid-Cretaceous of South America, eastern North America, and Africa. Mawsonia was first described by British paleontologist Arthur Smith Woodward in 1907.
Whiteia is an extinct genus of prehistoric coelacanth fish which lived during the Triassic period. It is named after Errol White.
Laugiidae is an extinct family of prehistoric marine coelacanths which lived during the Triassic and Jurassic periods. Their fossils have been found in Canada, Germany and Greenland.
Heptanema is an extinct genus of prehistoric coelacanth from the Middle Triassic (Ladinian) of northern Italy and southern Switzerland.
Hainbergia is an extinct genus of prehistoric sarcopterygian or lobe-finned fish.
Mylacanthus is an extinct genus of prehistoric coelacanth lobe-finned fish that lived during the Smithian age of the Early Triassic epoch in what is now Svalbard.
Sassenia is an extinct genus of prehistoric coelacanth lobe-finned fish that lived during the Early Triassic epoch in what is now East Greenland and Svalbard.
Acentrophorus is an extinct genus of prehistoric freshwater and marine ray-finned fish from the Roadian to the Wuchiapingian of England, Germany (Kupferschiefer), Italy and Russia. There may also be a Triassic occurrence in Australia.
Urosthenes is an extinct genus of prehistoric bony fish that lived during the Lopingian to Middle Triassic epochs in what is now New South Wales, Australia.
Mikko's Phylogeny Archive on Coelacanthiformes