Ticinepomis Temporal range: Lower Ladinian, | |
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Fossil | |
Reconstruction | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Sarcopterygii |
Class: | Actinistia |
Order: | Coelacanthiformes |
Family: | Latimeriidae |
Genus: | † Ticinepomis Rieppel, 1980 |
Type species | |
†Ticinepomis peyeri Rieppel, 1980 | |
Other species | |
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Ticinepomis is an extinct genus of coelacanth lobe-finned fish which lived during the Middle Triassic period in what is now Switzerland. It contains two species, T. peyeri and T. ducanensis. [2]
Specimens of the species T. peyeri, which was named after Bernhard Peyer, are most common in the Besano Formation (or Grenzbitumenzone) of Monte San Giorgio in canton Ticino. [3] Other coelacanths from Monte San Giorgio include a larger species (tentatively referred to Holophagus picenus ) from the Besano Formation, [4] and a species of Heptanema from the Meride Limestone. [5]
Larger Ticinepomis specimens have been found in the Prosanto Formation of canton Graubünden, originally referred to as Ticinepomis cf. T. peyeri. [6] A revision showed that this material belongs to a new species, T. ducanensis, remains of which were also discovered in the Besano Formation of canton Ticino. [2] The Prosanto Formation also produced the unusual coelacanths Foreyia and Rieppelia .
Ticinepomis was originally described as being a member of the family Coelacanthidae, being similar to Coelacanthus , Holophagus and Undina . [3] Later, T. peyeri was placed in Latimeriidae. [6] The bizarre Prosanto Formation latimeriid Foreyia is thought to be T. peyeri's closest relative, as they share many features despite their drastically contrasting appearances. [7]
Cymbospondylus is an extinct genus of large ichthyosaurs, of which it is among the oldest representatives, that lived from the Lower to Middle Triassic in what are now North America and Europe. The first known fossils of this taxon are a set of more or less complete vertebrae which were discovered in the 19th century in various mountain ranges of Nevada, in the United States, before being named and described by Joseph Leidy in 1868. It is in the beginning of the 20th century that more complete fossils were discovered through several expeditions launched by the University of California, and described in more detail by John Campbell Merriam in 1908, thus visualizing the overall anatomy of the animal. While many species have been assigned to the genus, only five are recognized as valid, the others being considered synonymous, doubtful or belonging to other genera. Cymbospondylus was formerly classified as a representative of the Shastasauridae, but more recent studies consider it to be more basal, view as the type genus of the Cymbospondylidae.
Ceresiosaurus is an extinct aquatic genus of lariosaurine nothosaurid sauropterygian known from the Middle Triassic of Monte San Giorgio, southern Switzerland and northern Italy. Ceresiosaurus, meaning "Lizard of Ceresio". The type species, Ceresiosaurus calcagnii, was named by Bernhard Peyer in 1931. C. calcagnii is known from both the Cava superiore and Cava inferiore beds of the Lower Meride Limestone at Monte San Giorgio, dating to the latest Anisian of the Middle Triassic. Rieppel (1998) suggested that the back then monospecific genus Ceresiosaurus, is a junior synonym of the better known Lariosaurus, yet he kept it type species as a separate species under the new combination L. calcagnii. In 2004, however, this synonymy was objected by Hänni who described and name a second species of Ceresiosaurus, C. lanzi - a separation supported by several other authors since. This species is known only from the stratigraphically younger Cassima beds of Monte San Giorgio, although also from the Lower Meride Limestone, dating to possibly the lowest Ladinian age. The species in a subtropical lagoonal environment with varying open marine influences, and alongside many related but smaller species of nothosaurids and pachypleurosaurids. Ceresiosaurus represents one of the largest vertebrate of up to 3 m (9.8 ft) snout-tail length from the very diversified paleoenvironment of the Middle Triassic Monte San Giorgio.
Coelacanthus is a genus of extinct marine coelacanths known from the late Permian period. It was the first genus of coelacanths described, about a century before the discovery of the extant coelacanth Latimeria. The order Coelacanthiformes is named after it.
Monte San Giorgio is a Swiss mountain and UNESCO World Heritage Site near the border between Switzerland and Italy. It is part of the Lugano Prealps, overlooking Lake Lugano in the Swiss Canton of Ticino.
Macrocnemus is an extinct genus of archosauromorph reptile known from the Middle Triassic of Europe and China. Macrocnemus is a member of the Tanystropheidae family and includes three species. Macrocnemus bassanii, the first species to be named and described, is known from the Besano Formation and adjacent paleontological sites in the Italian and Swiss Alps. Macrocnemus fuyuanensis, on the other hand, is known from the Zhuganpo Formation in southern China. A third species, Macrocnemus obristi, is known from the Prosanto Formation of Switzerland and is characterized by gracile limbs. The name Macrocnemus is Greek for "long tibia".
Holophagus is an extinct genus of coelacanth belonging to Latimeriidae. The type species, Holophagus gulo, is known from the Lower Jurassic marine Lias of England. Some authors have considered the genus restricted to the Lias of England.
Besanosaurus is a extinct genus of Middle Triassic ichthyosaur from Monte San Giorgio of Italy and Switzerland, containing the single species B. leptorhynchus. Besanosaurus was named by Cristiano Dal Sasso and Giovanni Pinna in 1996, based on the nearly complete flattened skeleton BES SC 999, the holotype specimen. This skeleton is preserved across multiple thin rock slabs spanning 3.5 by 4 metres when assembled and took thousands of hours to prepare. Additional specimens from Monte San Giorgio that have previously been considered separate genera, including a partial skull named Mikadocephalus and a well-preserved, largely complete skeleton, have been reinterpreted as additional specimens of Besanosaurus. Putative specimens of Besanosaurus have been discovered in the Norwegian archipelago of Svalbard and Germany, although their attribution to this genus remains disputed.
Serpianosaurus is an extinct genus of pachypleurosaurs known from the Middle Triassic deposits of Switzerland and Germany. It was a small reptile, with the type specimen of S. mirigiolensis measuring 75 cm (2.46 ft) long.
Besania is an extinct genus of prehistoric marine ray-finned fish that lived during the Anisian and Ladinian ages of the Middle Triassic epoch in what is now southern/southeastern Switzerland and northern Italy. Fossils were recovered from the Besano Formation of Monte San Giorgio area and the Prosanto Formation of canton Graubünden, Switzerland.
Ctenognathichthys is an extinct genus of prehistoric marine ray-finned fish that lived during the Middle Triassic epoch of Europe, in the former Tethys Ocean.
Heptanema is an extinct genus of prehistoric coelacanth from the Middle Triassic (Ladinian) of northern Italy and southern Switzerland.
Meridensia is an extinct genus of prehistoric ray-finned fish that lived during the Anisian and Ladinian ages of the Middle Triassic epoch in what is now southern Switzerland and northern Italy. Fossils were recovered from the Besano Formation of Monte San Giorgio and Besano area at the Swiss-Italian boundary.
Wimanius is a genus of ichthyosaur from the Middle Triassic of Switzerland, containing a single species, Wimanius odontopalatus. It was described by Michael Maisch and Andreas Matzke in 1998 based on an incomplete skull from Monte San Giorgio, a mountain on the Swiss-Italian border. Wimanius possesses teeth on its palate, though whether they were located on the palatine or pterygoid is disputed. Other features of Wimanius include a large orbit and jugals with two rami of similar lengths. Different phylogenetic placements of Wimanius have been recovered by different studies, including it being a mixosaurid relative or a merriamosaur, and a monotypic family, Wimaniidae has been named for it. However, its validity has also been questioned, and synonymy with various other genera has been proposed. The only specimen of Wimanius come from the Besano Formation. During the Anisian, this region was a lagoon populated by a wide variety of marine life, including a variety of other ichthyosaurs.
Eusaurosphargis is an extinct genus of a diapsid reptile, known from the Middle Triassic Besano Formation of northern Italy and Prosanto Formation of south-eastern Switzerland. It contains a single species, Eusaurosphargis dalsassoi. It was a small reptile, measuring 20 cm (7.9 in) long.
Helveticosauridae is an extinct family of basal marine reptiles known from the Middle Triassic of southern Switzerland and northern Italy.
Foreyia is an extinct genus of coelacanth lobe-finned fish which lived during the Middle Triassic period in what is now Canton of Graubünden, Switzerland. It contains a single species F. maxkuhni.
The Besano Formation is a geological formation in the southern Alps of northwestern Italy and southern Switzerland. This formation, a thin but fossiliferous succession of dolomite and black shale, is famous for its preservation of Middle Triassic (Anisian–Ladinian) marine life including fish and aquatic reptiles. It is exposed in the Monte San Giorgio and Besano area. It is among the formations responsible for the area being designated as a UNESCO World Heritage Site. In Switzerland, it is also known as the Grenzbitumenzone. The Anisian-Ladinian boundary lies in the upper part of the Besano Formation.
The San Salvatore Dolomite, sometimes known as the Salvatore Dolomite or San Salvatore Formation, is a Middle Triassic geological formation in Switzerland and Italy. The primarily lithology is micritic dolomite with a high proportion of algal mounds (stromatolites). It corresponds to a thick warm-water carbonate platform on the northern edge of an island in what is now the Po Plain. This formation and its local equivalents are common in the hills around Lake Maggiore, Varese, and Lugano, preserving fossils of marine invertebrates such as ammonoids, gastropods, and bivalves. At its southernmost extent on Monte San Giorgio, only the lower part of the San Salvatore Dolomite is preserved. The middle and upper parts are replaced by the Besano Formation, San Giorgio Dolomite, and Meride Limestone, which were deposited in a deeper and more anoxic basin between carbonate platforms.
Bernhard Peyer was a Swiss paleontologist and anatomist who served as a professor at the University of Zurich. A major contribution was on the evolution of vertebrate teeth.
Rieppelia is an extinct genus of latimeriid coelacanth fish from the Middle Triassic of Switzerland, in what is now Monte San Giorgio. It contains a single species, R. heinzfurreri.