Dominickus Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Lepidoptera |
Family: | Castniidae |
Genus: | † Dominickus Tindale, 1985 |
Species: | †D. castnioides |
Binomial name | |
†Dominickus castnioides Tindale, 1985 | |
Dominickus is an extinct genus of moth in the butterfly-moth family Castniidae containing a single species Dominickus castnioides. [1] The species is known from late Eocene, Priabonian stage, [2] lake deposits near the small community of Guffey in Teller County, Colorado, United States. [3]
Dominickus castnioides is known only from one fossil, the holotype, specimen "P.22949". It is a single, mostly complete pair of fore wings, preserved as a compression fossil in fine grained shale. [1] The shale specimen is from the fossiliferous outcrops of the Florissant Formation which outcrop near Guffey 10 miles (16 km) southwest of Florissant. The type specimen is currently preserved in the paleoentomological collections housed in the Field Museum of Natural History, located in Chicago, Illinois, United States. Dominickus was first studied by Norman Barnett Tindale of Palo Alto, California, with his 1985 type description being published in the Journal of Research on the Lepidoptera. [1] The generic name was coined by Tindale in honor of Richard B. Dominick. Tindale did not provide an explanation for the specific epithet castnioides.
When first examined, the specimen was thought to possibly be related to an early ancestor of Hesperioidea and Papilionoidea, due to the overall wide shape of the wings and the fork of the Cu vein being near the wing midpoint. [1] However while Tindale was examining a series of photographs taken in Australia of live Synemon species butterfly-moths, he noticed a strong similarity between the modern moth and the Florissant fossil. [1] Tindale then compared the fossil to the modern Chilean species Castnia psittacus which shows a closer match. The major difference between the fossil and modern members of Castniidae is the fossil's lack of a crossvein between the R1 and R2 veins. [1]
At the time of description, the Florissant Formation was considered to be Oligocene in age. [1] Further refinement of the formation's age using radiometric dating of sanidine crystals has resulted in an age of 34 million years old, which places the formation in the Eocene Chadronian stage. [4] [5] [6] Further refinement of the Eocene-Oligocene has placed the formation into the late Eocene Priabonian stage. [2]
The holotype forewings of Dominickus castnioides are about 16.4 millimetres (0.65 in) long and about 9.4 millimetres (0.37 in) wide. The wings have an overall broadly triangular shape with a rounded tip. The vein structure is very similar to that of modern Castniidae members, having an R-vein which supports the costal third of the wing. Due to the nature of the matrix in which the fossil is preserved, the wing scales of the specimen are not distinguishable or are absent. The outline of wing margin furthest from the wing base is possibly a fringe like that found in the modern members of the family. Photographs of the specimen give possible indications of the color pattering with the costal area, narrow at the wing base and widening out at the midpoint a dark tone. The base of the wing was possibly light colored with the tone extending along the hind margin. [1]
The Florissant Formation is a sedimentary geologic formation outcropping around Florissant, Teller County, Colorado. The formation is noted for the abundant and exceptionally preserved insect and plant fossils that are found in the mudstones and shales. Based on argon radiometric dating, the formation is Eocene in age and has been interpreted as a lake environment. The fossils have been preserved because of the interaction of the volcanic ash from the nearby Thirtynine Mile volcanic field with diatoms in the lake, causing a diatom bloom. As the diatoms fell to the bottom of the lake, any plants or animals that had recently died were preserved by the diatom falls. Fine layers of clays and muds interspersed with layers of ash form "paper shales" holding beautifully-preserved fossils. The Florissant Fossil Beds National Monument is a national monument established to preserve and study the geology and history of the area.
Florissantia is an extinct genus of flowering plants in the Malvaceae subfamily Sterculioideae known from western North America and far eastern Asia. Flower, fruit, and pollen compression fossils have been found in formations ranging from the Early Eocene through to the Early Oligocene periods. The type species is Florissantia speirii and three additional species are known, Florissantia ashwillii, Florissantia quilchenensis, and Florissantia sikhote-alinensis.
Florissantoraphidia is an extinct genus of snakefly in the family Raphidiidae containing two described species Florissantoraphidia funerata and Florissantoraphidia mortua. Both species were originally placed in the living raphidiid genus Raphidia, as Raphidia funerata and Raphidia mortua respectively. before being redescribed and transferred to the newly erected genus in 2014.
Prodryas persephone is an extinct species of brush-footed butterfly, known from a single specimen from the Chadronian-aged Florissant Shale Lagerstätte of Late Eocene Colorado. P. persephone is the first fossil butterfly to be found in North America, and is exquisitely well preserved. Its closest extant relatives are the mapwings and African admirals of the genera Hypanartia and Antanartia, respectively.
Palaeovespa is an extinct genus of wasp in the Vespidae subfamily Vespinae. The genus currently contains eight species: five from the Priabonian stage Florissant Formation in Colorado, United States, two from the middle Eocene Baltic amber deposits of Europe, and one species from the late Paleocene of France.
Anthidium scudderi is an extinct species of mason bee in the Megachilidae genus Anthidium. The species is solely known from the late Eocene, Chadronian stage, Florissant Formation deposits in Florissant, Colorado, USA. Anthidium scudderi is one of only four extinct species of mason bees known from the fossil record, and with Anthidium exhumatum, one of two species from the Florissant Formation.
Anthidium exhumatum is an extinct species of mason bee in the Megachilidae genus Anthidium. The species is solely known from the late Eocene, Chadronian stage, Florissant Formation deposits in Florissant, Colorado, USA. Anthidium exhumatum is one of only four extinct species of mason bees known from the fossil record, and with Anthidium scudderi, one of two species from the Florissant Formation.
The Lepidoptera fossil record encompasses all butterflies and moths that lived before recorded history. The fossil record for Lepidoptera is lacking in comparison to other winged species, and tending not to be as common as some other insects in the habitats that are most conducive to fossilization, such as lakes and ponds, and their juvenile stage has only the head capsule as a hard part that might be preserved. Yet there are fossils, some preserved in amber and some in very fine sediments. Leaf mines are also seen in fossil leaves, although the interpretation of them is tricky. Putative fossil stem group representatives of Amphiesmenoptera are known from the Triassic.
Paleolepidopterites is a collective genus of fossil moths which can not be placed in any defined family. The included species were formerly placed in the leaf-roller family Tortricidae and are known from fossils found in Russia and the United States. The collective genus contains three species: Paleolepidopterites destructus, Paleolepidopterites florissantanus, and Paleolepidopterites sadilenkoi, formerly placed within the genera Tortrix and Tortricites respectively. The three species were formally redescribed and moved to the new collective genus by Heikkilä et al. (2018).
Hydriomena? protrita is an extinct species of moth in the family Geometridae, and possibly in the modern genus Hydriomena. The species is known from late Eocene, Priabonian stage, lake deposits of the Florissant Formation in Teller County, Colorado, United States. It was first described by Theodore Dru Alison Cockerell in 1922.
Eulithomyrmex is an extinct genus of ant in the formicid subfamily Agroecomyrmecinae. The genus contains two described species, Eulithomyrmex rugosus and Eulithomyrmex striatus. Eulithomyrmex is known from a group of Late Eocene fossils which were found in North America.
Aphaenogaster donisthorpei is an extinct species of ant in formicid subfamily Myrmicinae known from a Late Eocene fossil from North America. A. donisthorpei was one of two Aphaenogaster species described in the 1930 paper.
The White River Formation is a geologic formation of the Paleogene Period, in the northern Great Plains and central Rocky Mountains, within the United States.
Aphaenogaster mayri is an extinct species of ant in formicid subfamily Myrmicinae known from a series of Late Eocene fossils found in North America. A. mayri was one of two Aphaenogaster species described in a 1930 paper by Frank M. Carpenter.
Archiponera is an extinct genus of ant in the formicid subfamily Ponerinae. The genus contains a single described species, Archiponera wheeleri known from several Late Eocene fossils which were found in North America.
Priabona is an extinct genus of big-headed flies in the dipteran subfamily Nephrocerinae, within which it is one of only two genera. The genus contains a single described species, Priabona florissantius. Priabona is known from a single Late Eocene fossil from western North America.
Holcorpa is a genus of extinct insects in the scorpionfly order Mecoptera. Two Eocene age species found in Western North America were placed into the genus, H. dillhoffi and H. maculosa.
Florissantia is an extinct monotypic genus of planthopper in the dictyopharid subfamily Dictyopharinae. The single species, Florissantia elegans, was described by Samuel Hubbard Scudder (1890) from fossils found in the Florissant Formation of Colorado.
Dipteronia brownii is an extinct species in the soapberry family (Sapindaceae) described in 2001. Fossils of D. brownii are known from stratigraphic formations in North America and Asia ranging in age between Paleocene to Early Oligocene.
Polystoechotites is an extinct parataxon of lacewings in the moth lacewing family Ithonidae. The taxon is a collective group for fossil polystechotid giant lacewing species whose genus affiliation is uncertain, but which are distinct enough to identify as segregate species. Polystoechotites species are known from Eocene fossils found in North America and is composed of four named species Polystoechotites barksdalae, Polystoechotites falcatus, Polystoechotites lewisi, and Polystoechotites piperatus, plus two unnamed species. Three of the described species are known from fossils recovered from the Eocene Okanagan Highlands of Washington State, while the fourth is from Colorado.