A common classification of the Lepidoptera involves their differentiation into butterflies and moths. Butterflies are a natural monophyletic group, often given the suborder Rhopalocera, which includes Papilionoidea, Hesperiidae (skippers), and Hedylidae. In this taxonomic scheme, moths belong to the suborder Heterocera. Other taxonomic schemes have been proposed, the most common putting the butterflies into the suborder Ditrysia and then the "superfamily" Papilionoidea and ignoring a classification for moths.
The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a reduced pair of forelegs and many hold their colourful wings flat when resting. They are also called brush-footed butterflies or four-footed butterflies, because they are known to stand on only four legs while the other two are curled up; in some species, these forelegs have a brush-like set of hairs, which gives this family its other common name. Many species are brightly coloured and include popular species such as the emperors, monarch butterfly, admirals, tortoiseshells, and fritillaries. However, the under wings are, in contrast, often dull and in some species look remarkably like dead leaves, or are much paler, producing a cryptic effect that helps the butterflies blend into their surroundings.
Skippers are a group of butterflies placed in the family Hesperiidae within the order Lepidoptera. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy places the family in the superfamily Papilionoidea, the butterflies. They are named for their quick, darting flight habits. Most have their antenna tips modified into narrow, hook-like projections. Moreover, skippers mostly have an absence of wing-coupling structure available in most moths. More than 3500 species of skippers are recognized, and they occur worldwide, but with the greatest diversity in the Neotropical regions of Central and South America.
Lycaenidae is the second-largest family of butterflies, with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfly species.
Riodinidae is the family of metalmark butterflies. The common name "metalmarks" refers to the small, metallic-looking spots commonly found on their wings. The 1,532 species are placed in 146 genera. Although mostly Neotropical in distribution, the family is also represented both in the Nearctic, Palearctic, Australasian (Dicallaneura), Afrotropic, and Indomalayan realms.
The Nymphalinae are a subfamily of brush-footed butterflies. Sometimes, the subfamilies Limenitidinae, and Biblidinae are included here as subordinate tribe(s), while the tribe Melitaeini is occasionally regarded as a distinct subfamily.
Hamearis lucina, the Duke of Burgundy, the only member of the genus Hamearis, is a European butterfly in the family Riodinidae. For many years, it was known as the "Duke of Burgundy fritillary", because the adult's chequered pattern is strongly reminiscent of "true" fritillaries of the family Nymphalidae.
The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae. They were formerly considered a distinct family, Satyridae. This group contains nearly half of the known diversity of brush-footed butterflies. The true number of the Satyrinae species is estimated to exceed 2,400.
The Morphinae are a subfamily of Nymphalidae butterflies that includes the morphos, the owl butterflies (Caligo), and related lineages. It is either considered a sister group of the Satyrinae, or disassembled and included therein.
Hedylidae, the "American moth-butterflies", is a family of insects in the order Lepidoptera, representing the superfamily Hedyloidea. They have traditionally been viewed as an extant sister group of the butterfly superfamily Papilionoidea, but a 2014 phylogenetic analyses has suggested Hedylidae is a subgroup of Papilionoidea, and not a sister group, and are more accurately referred to as butterflies rather than moths. They are represented by a single Neotropical genus Macrosoma with 35 currently recognized species.
Butterfly evolution is the origin and diversification of butterflies through geologic time and over a large portion of the Earth's surface. The earliest known butterfly fossils are from the mid Eocene epoch, between 40-50 million years ago. Their development is closely linked to the evolution of flowering plants, since both adult butterflies and caterpillars feed on flowering plants. Of the 220,000 species of Lepidoptera, about 45,000 species are butterflies, which probably evolved from moths. Butterflies are found throughout the world, except in Antarctica, and are especially numerous in the tropics; they fall into eight different families.
The Elymniini is one of the tribes of the subfamily Satyrinae. If the subfamily Satyrinae elevate to family status, this tribe shall be treated as subfamily Elymniinae. Elymniini was formerly a large group, but recently, it is considered to be include only one genus, Elymnias, according to molecular phylogenetic analyses.
Brassolini is a tribe usually placed in the brush-footed butterfly subfamily Morphinae, which is often included in the Satyrinae as a tribe Morphini. If this is accepted, the Brassolini become the sister tribe of the Morphini among the Satyrinae. Formerly, they were treated as an independent family Brassolidae or subfamily Brassolinae. Many members of this tribe are called owl butterflies.
Hesperocharis is a genus of butterflies in the family Pieridae. They are native to the Americas.
Taygetis is a genus of satyrid butterflies found in the Neotropical realm.
Hyantis is a monotypic nymphalid butterfly genus. Its sole species is Hyantis hodeva, which is found in New Guinea. It is uncertain which tribe this butterfly should be placed within.
Styx is a monotypic genus of butterflies in the metalmark family Riodinidae. It consists of one species, Styx infernalis, described by Otto Staudinger in 1875. It is endemic to Peru, where it inhabits tropical montane cloud forests between the elevations of 1000-1600 meters.
