Many-plumed moths | |
---|---|
Twenty-plume moth (Alucita hexadactyla: Alucitidae) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Lepidoptera |
Superfamily: | Alucitoidea |
Family: | Alucitidae Leach, 1815 |
Type species | |
Alucita hexadactyla | |
Diversity | |
9 genera, about 210 species | |
Synonyms | |
Alucitina Zeller, 1841 |
The Alucitidae or many-plumed moths [1] are a family of moths with unusually modified wings. Both fore- and hind-wings consist of about six rigid spines, from which radiate flexible bristles creating a structure similar to a bird's feather.
This is a small family, with about a global total of 210 species described to date (though it is likely that some undescribed species remain to be discovered). They are found mostly in temperate to subtropical (but not tropical) regions.
The taxonomy of this family is somewhat disputed. Here, they are united in superfamily Alucitoidea with the Tineodidae, a diverse group of numerous small genera with about 20 species all together. However, the two supposed Alucitoidea families may be polyphyletic with regard to each other, and Tineodidae better included in Alucitidae. In any case, the similar-looking plume moths (Pterophoroidea) are widely held to be very close, if not the closest living relatives of the Alucitoidea. [2]
Earlier, many authors assumed that the fruitworm moths (Copromorphoidea) were also very closely related to the Alucitidae (and the fringe-tufted moths, Epermeniidae) – according to some, closer in fact than the Pterophoroidea and even the Tineodidae. In this Alucitoidea do not exist; Alucitidae and Tineodidae are assigned to different (but still most closely related) superfamilies. In the treatment here, the Copromorphoidea are presumed to be the most advanced of these lineages of small but fairly "modern" moths, while the Alucitoidea and Pterophoroidea are more primitive. [2] [3]
The genera presently placed here, sorted alphabetically, are: [3]
Oecophoridae is a family of small moths in the superfamily Gelechioidea. The phylogeny and systematics of gelechoid moths are still not fully resolved, and the circumscription of the Oecophoridae is strongly affected by this.
The Elachistidae are a family of small moths in the superfamily Gelechioidea. Some authors lump about 3,300 species in eight subfamilies here, but this arrangement almost certainly results in a massively paraphyletic and completely unnatural assemblage, united merely by symplesiomorphies retained from the first gelechioid moths.
The Pyralidae, commonly called pyralid moths, snout moths or grass moths, are a family of Lepidoptera in the ditrysian superfamily Pyraloidea. In many classifications, the grass moths (Crambidae) are included in the Pyralidae as a subfamily, making the combined group one of the largest families in the Lepidoptera. The latest review by Eugene G. Munroe and Maria Alma Solis retain the Crambidae as a full family of Pyraloidea.
The Pyraloidea are a moth superfamily containing about 16,000 described species worldwide, and probably at least as many more remain to be described. They are generally fairly small moths, and as such, they have been traditionally associated with the paraphyletic Microlepidoptera.
Gelechioidea is the superfamily of moths that contains the case-bearers, twirler moths, and relatives, also simply called curved-horn moths or gelechioid moths. It is a large and poorly understood '"micromoth" superfamily, constituting one of the basal lineages of the Ditrysia.
Aluctoidea is the superfamily of many-plumed and false plume moths. These small moths are most easily recognized by their wings. These each consist of many narrow strips of membrane around the major veins, instead of a continuous sheet of membrane between the veins. In living moths in the wild, this is often hard to see however. When they are at rest, the "plumes" partly overlap, appearing as solid wings. But even then, they can be recognized by the wings having a marked lengthwise pattern and uneven edge.
The Coleophoridae are a family of small moths, belonging to the huge superfamily Gelechioidea. Collectively known as case-bearers, casebearing moths or case moths, this family is represented on all continents, but the majority are found in temperate areas of the Northern Hemisphere. They are most common in the Palearctic, and rare in sub-Saharan Africa, South America, and Australia; consequently, they probably originated in northern Eurasia. They are relatively common in houses, they seek out moist areas to rest and procreate.
Callidulidae, the only known family of the superfamily Calliduloidea, is the family of Old World butterfly-moths, containing eight genera. They have a peculiar distribution, restricted to the Old World tropics of Southeast Asia to Australasia and Madagascar. The three subfamilies exhibit both day- and night-flying behaviour.
Choreutidae, or metalmark moths, are a family of insects in the lepidopteran order whose relationships have been long disputed. It was placed previously in the superfamily Yponomeutoidea in family Glyphipterigidae and in superfamily Sesioidea. It is now considered to represent its own superfamily. The relationship of the family to the other lineages in the group "Apoditrysia" need a new assessment, especially with new molecular data.
Epermeniidae or the fringe-tufted moths is a family of insects in the lepidopteran order with about 14 genera. Previously they have been divided in two subfamilies Epermeniinae and Ochromolopinae but this is no longer maintained since the last group is probably hierarchically nested within the first. They are presently placed in their own superfamily but have previously been placed among the Yponomeutoidea or Copromorphoidea with which they share some features. Their systematic placement among the apoditrysian group "Obtectomera" is however uncertain. They show some morphological similarities to the "plume moths", for example the wing fringe has similar groups of scales. There are also some similarities to Schreckensteinioidea, for example spiny legs and at least in some species an open-network cocoon. The genus Thambotricha from New Zealand may be the sister group of all other extant members. The most important genera are Epermenia, Ochromolopis and Gnathifera. The group has been extensively revised and catalogued by Dr Reinhard Gaedike.
Carposinoidea, the "fruitworm moths", is a superfamily of insects in the lepidopteran order. The superfamily is also known as Copromorphoidea, which is a junior synonym. These moths are small to medium-sized and are broad-winged bearing some resemblance to the superfamilies Tortricoidea and Immoidea. The antennae are often "pectinate" especially in males, and many species of these well camouflaged moths bear raised tufts of scales on the wings and a specialised fringe of scales at the base of the hindwing sometimes in females only; there are a number of other structural characteristics. The position of this superfamily is not certain, but it has been placed in the natural group of "Apoditrysia" "Obtectomera", rather than with the superfamilies Alucitoidea or Epermenioidea within which it has sometimes previously been placed, on the grounds that shared larval and pupal characteristics of these groups have probably evolved independently. It has been suggested that the division into two families should be abandoned.
The Tineodidae or false plume moths are a family of moths with in some cases unusually modified wings: Like in some related moths, the wings of several Tineodidae are decomposed into several rigid spines. This is a small family, with about a global total of 20 species described to date; some undescribed species are known or suspected to exist however. They seem to be of Australian origin, where they are most diverse, but range through the Wallacea to Southeast and South Asia, and into the Pacific to the Marquesas Islands.
Sematuridae is a family of moths in the lepidopteran order that contains two subfamilies.
The insect order Lepidoptera consists of moths, most of which are night-flying, and a derived group, mainly day-flying, called butterflies. Within Lepidoptera as a whole, the groups listed below before Glossata contain a few basal families accounting for less than 200 species; the bulk of Lepidoptera are in the Glossata. Similarly, within the Glossata, there are a few basal groups listed first, with the bulk of species in the Heteroneura. Basal groups within Heteroneura cannot be defined with as much confidence, as there are still some disputes concerning the proper relations among these groups. At the family level, however, most groups are well defined, and the families are commonly used by hobbyists and scientists alike.
The Obtectomera is a clade of macro-moths and butterflies, comprising over 100,000 species in at least 12 superfamilies.
Alucita is the largest genus of many-plumed moths ; it is also the type genus of its family and the disputed superfamily Alucitoidea. This genus occurs almost worldwide and contains about 180 species as of 2011; new species are still being described and discovered regularly. Formerly, many similar moths of superfamilies Alucitoidea, Copromorphoidea and Pterophoroidea were also placed in Alucita.
Triscaedecia is a genus of moths in the family Alucitidae, found in the Malay and Polynesian regions. Triscaedecia is distinguished from all other genera in the family by the hindwings being split into seven, instead of six, lobes. The lobes in both wings are only split to about two-thirds of the length of the wings. The genus was erected by George Hampson in 1905.
Epimarptidae was a former, or is a possible, monotypic family of moths in the moth superfamily Gelechioidea. It can now be seen as either a synonym of family Batrachedridae, or a monotypic subfamily of that family.