Cossidae

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Cossidae
Cossulamagnifica.JPG
Cossula magnifica (Cossulinae)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Infraorder: Heteroneura
Clade: Eulepidoptera
Clade: Ditrysia
Clade: Apoditrysia
Superfamily: Cossoidea
Family: Cossidae
Leach, 1815

The Cossidae, the cossid millers or carpenter millers, make up a family of mostly large miller moths. This family contains over 110 genera with almost 700 known species, and many more species await description. Carpenter millers are Lepidoptera found worldwide, They are nocturnal, except for the Southeast Asian subfamily Ratardinae.

Contents

Witchetty grubs (Endoxyla leucomochla) of the subfamily Zeuzerinae Witchetty grub.jpg
Witchetty grubs ( Endoxyla leucomochla ) of the subfamily Zeuzerinae

This family includes many species with large caterpillars and moths with a wingspan from 9–24 cm (3+129+12 in).[ citation needed ] These moths are mostly grey; some have long, narrow wings and resemble hawkmoths (Sphingidae) which are more advanced macrolepidoptera, however. Many are twig, bark, or leaf mimics, and Cossidae often have some sort of large marking at the tip of the forewing uppersides, conspicuous in flight, but resembling a broken-off twig when the animals are resting.

Caterpillars are smooth with a few hairs. [1] Most cossid caterpillars are tree borers, in some species taking up to three years to mature. The caterpillars pupate within their tunnels; they often have an unpleasant smell, hence another colloquial name is goat moths.

The family includes the carpenterworm ( Prionoxystus robiniae ) and the goat moth (Cossus cossus) which have gained popularity as pests. However, the large caterpillars of species that do not smell badly are often edible. Witchetty grubs among the Outback's most famous bush tucker are most commonly the caterpillars of Endoxyla leucomochla , one of the more than 80 cossid species in Australia. In Chile, the sweet-smelling caterpillars of the Chilean moth ( Chilecomadia moorei ) are harvested in quantity and internationally traded as butterworms, for use as pet food and fishing bait.

Systematics

Some other animal families, such as the Dudgeoneidae, Metarbelidae, and Ratardidae, have been included within this family time and again. The first is considered a distinct family of the Cossoidea today recognizable by their abdominal tympanal organs which the Cossidae lack, whereas the other two are usually kept in the Cossidae as subfamilies. Some unrelated millers were included in the Cossidae in error, too, such as the genus Holcoceroides which is more primitive Ditrysia, or the Andesianidae which are even more ancient Heteroneura.

The Cossidae were usually divided into six subfamilies. [2] However, numerous new subfamilies have been described recently, the current taxonomy is:

Specimen of the large Zeuzerinae genus Xyleutes Drevotochets evkaliptovyi.jpg
Specimen of the large Zeuzerinae genus Xyleutes
Zyganisus caliginosus belongs to an Australian genus of unclear affiliations. Zyganisus caliginosus (ento-csiro-au).jpg
Zyganisus caliginosus belongs to an Australian genus of unclear affiliations.

Excluded genera

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<span class="mw-page-title-main">Pyralidae</span> Family of moths

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<span class="mw-page-title-main">Coleophoridae</span> Family of moths

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<span class="mw-page-title-main">Cossinae</span> Subfamily of moths

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<span class="mw-page-title-main">Nacophorini</span> Tribe of moths

The Nacophorini are one of the smaller tribes of geometer moths in the subfamily Ennominae. They are the most diverse Ennominae of Australia and are widespread in the Americas. If the African genera tentatively placed herein indeed belong here, the distribution of the Nacophorini is distinctly Gondwanan, with their probable origin either of Australia, South America or even Antarctica. In Eurasia, they are rare by comparison.

<i>Cossus cossus</i> Species of moth

Cossus cossus, the goat moth, is a moth of the family Cossidae. It is found in Northern Africa, Asia and Europe.

<i>Nemapogon granella</i> Species of moth

Nemapogon granella is a species of tineoid moth. It belongs to the fungus moth family (Tineidae), and therein to the subfamily Nemapogoninae. It is the type species of its genus Nemapogon, and via that also of the subfamily Nemapogoninae. It is also the type species of the proposed genera Brosis and Diaphthirusa, which are consequently junior objective synonyms of Nemapogon.

<i>Oecophora bractella</i> Species of moth

Oecophora bractella is a species of gelechioid moth. It belongs to the subfamily Oecophorinae of the concealer moth family (Oecophoridae). As the type species of its genus Oecophora, its affiliations and phylogeny determine the delimitation of that family and subfamily.

<i>Borkhausenia minutella</i> Species of moth

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<i>Alabonia geoffrella</i> Species of moth

Alabonia geoffrella is a species of gelechioid moth. Here, it is placed within the subfamily Oecophorinae of the concealer moth family (Oecophoridae). Alternatively it has been placed in the Elachistidae or Depressariinae together with its presumed closest relatives.

<i>Eugraphe</i> Genus of moths

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<i>Nemaxera</i> Genus of moths

Nemaxera is a genus of the fungus moth family, Tineidae. Therein, it belongs to the subfamily Nemapogoninae. The genus is considered monotypic, with the single species Nemaxera betulinella placed here.

<i>Scythris</i> Genus of moths

Scythris is a genus of gelechioid moths. It is the type genus of the flower moth family, which is sometimes included as a subfamily in the Xyloryctidae, or together with these merged into the Oecophoridae. The genus was erected by Jacob Hübner in 1825.

<span class="mw-page-title-main">Galleriinae</span> Subfamily of moths

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<i>Cossus</i> Genus of moths

Cossus is a genus of moths in the family Cossidae described by Johan Christian Fabricius in 1793.

Cossus dentilinea is a moth in the family Cossidae first described by Herbert Druce in 1911. It is found in Colombia.

<span class="mw-page-title-main">Metarbelidae</span> Family of moths

The Metarbelidae are a family of the Cossoidea also called the carpenter or goat moths, and is sometimes treated as a subfamily, Metarbelinae of the Cossidae. No synapomorphies are shared with the Cossidae based on adult morphology. The family Metarbelidae was first described by Embrik Strand in 1909.

References

  1. Hampson, G. F. (1892). The Fauna of British India, Including Ceylon and Burma: Moths Volume I. Taylor and Francis. Archived from the original on 2017-09-30. Retrieved 2018-01-14 via Biodiversity Heritage Library.
  2. Pitkin & Jenkins (2004) and see references in Savela (2006)

Further reading