Cossidae | |
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Cossula magnifica (Cossulinae) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Lepidoptera |
Infraorder: | Heteroneura |
Clade: | Eulepidoptera |
Clade: | Ditrysia |
Clade: | Apoditrysia |
Superfamily: | Cossoidea |
Family: | Cossidae Leach, 1815 |
The Cossidae, the cossid millers or carpenter millers, make up a family of mostly large miller moths. This family contains over 110 genera with almost 700 known species, and many more species await description. Carpenter millers are Lepidoptera found worldwide, They are nocturnal, except for the Southeast Asian subfamily Ratardinae.
This family includes many species with large caterpillars and moths with a wingspan from 9–24 cm (3+1⁄2–9+1⁄2 in).[ citation needed ] These moths are mostly grey; some have long, narrow wings and resemble hawkmoths (Sphingidae) which are more advanced macrolepidoptera, however. Many are twig, bark, or leaf mimics, and Cossidae often have some sort of large marking at the tip of the forewing uppersides, conspicuous in flight, but resembling a broken-off twig when the animals are resting.
Caterpillars are smooth with a few hairs. [1] Most cossid caterpillars are tree borers, in some species taking up to three years to mature. The caterpillars pupate within their tunnels; they often have an unpleasant smell, hence another colloquial name is goat moths.
The family includes the carpenterworm ( Prionoxystus robiniae ) and the goat moth (Cossus cossus) which have gained popularity as pests. However, the large caterpillars of species that do not smell badly are often edible. Witchetty grubs – among the Outback's most famous bush tucker – are most commonly the caterpillars of Endoxyla leucomochla , one of the more than 80 cossid species in Australia. In Chile, the sweet-smelling caterpillars of the Chilean moth ( Chilecomadia moorei ) are harvested in quantity and internationally traded as butterworms, for use as pet food and fishing bait.
Some other animal families, such as the Dudgeoneidae, Metarbelidae, and Ratardidae, have been included within this family time and again. The first is considered a distinct family of the Cossoidea today recognizable by their abdominal tympanal organs which the Cossidae lack, whereas the other two are usually kept in the Cossidae as subfamilies. Some unrelated millers were included in the Cossidae in error, too, such as the genus Holcoceroides which is more primitive Ditrysia, or the Andesianidae which are even more ancient Heteroneura.
The Cossidae were usually divided into six subfamilies. [2] However, numerous new subfamilies have been described recently, the current taxonomy is:
The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek geo γεω, and metron μέτρον "measure" in reference to the way their larvae, or inchworms, appear to measure the earth as they move along in a looping fashion. Geometridae is a very large family, containing around 23,000 described species; over 1400 species from six subfamilies are indigenous to North America alone. A well-known member is the peppered moth, Biston betularia, which has been the subject of numerous studies in population genetics. Several other geometer moths are notorious pests.
The Pyralidae, commonly called pyralid moths, snout moths or grass moths, are a family of Lepidoptera in the ditrysian superfamily Pyraloidea. In many classifications, the grass moths (Crambidae) are included in the Pyralidae as a subfamily, making the combined group one of the largest families in the Lepidoptera. The latest review by Eugene G. Munroe and Maria Alma Solis retain the Crambidae as a full family of Pyraloidea.
The Coleophoridae are a family of small moths, belonging to the huge superfamily Gelechioidea. Collectively known as case-bearers, casebearing moths or case moths, this family is represented on all continents, but the majority are found in temperate areas of the Northern Hemisphere. They are most common in the Palearctic, and rare in sub-Saharan Africa, South America, and Australia; consequently, they probably originated in northern Eurasia. They are relatively common in houses, they seek out moist areas to rest and procreate.
Xestia is a genus of noctuid moths. They are the type genus of the tribe Xestiini in subfamily Noctuinae, though some authors merge this tribe with the Noctuini. Species in this genus are commonly known as "clays", "darts" or "rustics", but such names are commonplace among Noctuidae. Xestia moths have a wide distribution, though they most prominently occur in the Holarctic.
The Blastobasidae are a family of moths in the superfamily Gelechioidea. Its species can be found almost anywhere in the world, though in some places they are not native but introduced by humans. In some arrangements, these moths are included in the case-bearer family (Coleophoridae) as subfamily Blastobasinae. The Symmocidae are sometimes included in the Blastobasidae as subfamily or tribe.
The Cossinae are the nominate subfamily of the Cossidae. The caterpillars of several Cossinae species, such as the carpenterworm and the goat moth, are significant pests. On the other hand, in Chile the caterpillars of the Chilean moth are collected on a commercial scale for sale as fishing bait and terrarium pet food; they are usually called "butterworms" in international trade.
The Boarmiini are a large tribe of geometer moths in the Ennominae subfamily.
The Nacophorini are one of the smaller tribes of geometer moths in the subfamily Ennominae. They are the most diverse Ennominae of Australia and are widespread in the Americas. If the African genera tentatively placed herein indeed belong here, the distribution of the Nacophorini is distinctly Gondwanan, with their probable origin either of Australia, South America or even Antarctica. In Eurasia, they are rare by comparison.
Cossus cossus, the goat moth, is a moth of the family Cossidae. It is found in Northern Africa, Asia and Europe.
Nemapogon granella is a species of tineoid moth. It belongs to the fungus moth family (Tineidae), and therein to the subfamily Nemapogoninae. It is the type species of its genus Nemapogon, and via that also of the subfamily Nemapogoninae. It is also the type species of the proposed genera Brosis and Diaphthirusa, which are consequently junior objective synonyms of Nemapogon.
Oecophora bractella is a species of gelechioid moth. It belongs to the subfamily Oecophorinae of the concealer moth family (Oecophoridae). As the type species of its genus Oecophora, its affiliations and phylogeny determine the delimitation of that family and subfamily.
Borkhausenia minutella is a species of moth. Within its superfamily, it is placed within the subfamily Oecophorinae of the "concealer moth" family, Oecophoridae.
Alabonia geoffrella is a species of gelechioid moth. Here, it is placed within the subfamily Oecophorinae of the concealer moth family (Oecophoridae). Alternatively it has been placed in the Elachistidae or Depressariinae together with its presumed closest relatives.
Eugraphe is a genus of noctuid moths. They belong to the tribe Xestiini of the typical noctuid subfamily Noctuinae, though some do not separate this tribe and include the genus in the Noctuini. It is closely related to Anagnorisma, Coenophila and Eugnorisma, and as it seems most closely to the first of these. The geographic range is Palearctic, north of the Alpides but including the Caucasus, and between the Arctic and the arid lands of Central Asia.
Nemaxera is a genus of the fungus moth family, Tineidae. Therein, it belongs to the subfamily Nemapogoninae. The genus is considered monotypic, with the single species Nemaxera betulinella placed here.
Scythris is a genus of gelechioid moths. It is the type genus of the flower moth family, which is sometimes included as a subfamily in the Xyloryctidae, or together with these merged into the Oecophoridae. The genus was erected by Jacob Hübner in 1825.
The Galleriinae are a subfamily of snout moths and occur essentially worldwide, in some cases aided by involuntary introduction by humans. This subfamily includes the wax moths, whose caterpillars (waxworms) are bred on a commercial scale as food for pets and as fishing bait; in the wild, these and other species of Galleriinae may also be harmful to humans as pests.
Cossus is a genus of moths in the family Cossidae described by Johan Christian Fabricius in 1793.
Cossus dentilinea is a moth in the family Cossidae first described by Herbert Druce in 1911. It is found in Colombia.
The Metarbelidae are a family of the Cossoidea also called the carpenter or goat moths, and is sometimes treated as a subfamily, Metarbelinae of the Cossidae. No synapomorphies are shared with the Cossidae based on adult morphology. The family Metarbelidae was first described by Embrik Strand in 1909.