Cossidae | |
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Cossula magnifica (Cossulinae) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Lepidoptera |
Infraorder: | Heteroneura |
Clade: | Eulepidoptera |
Clade: | Ditrysia |
Clade: | Apoditrysia |
Superfamily: | Cossoidea |
Family: | Cossidae Leach, 1815 |
The Cossidae, the cossid millers or carpenter millers, make up a family of mostly large miller moths. This family contains over 110 genera with almost 700 known species, and many more species await description. Carpenter millers are nocturnal Lepidoptera found worldwide, except the Southeast Asian subfamily Ratardinae, which is mostly active during the day.
This family includes many species with large caterpillars and moths with a wingspan from 9–24 cm (3+1⁄2–9+1⁄2 in).[ citation needed ] These moths are mostly grey; some have long, narrow wings and resemble hawkmoths (Sphingidae) which are more advanced macrolepidoptera, however. Many are twig, bark, or leaf mimics, and Cossidae often have some sort of large marking at the tip of the forewing uppersides, conspicuous in flight, but resembling a broken-off twig when the animals are resting.
Caterpillars are smooth with a few hairs. [1] Most cossid caterpillars are tree borers, in some species taking up to three years to mature. The caterpillars pupate within their tunnels; they often have an unpleasant smell, hence another colloquial name is goat moths.
The family includes the carpenterworm ( Prionoxystus robiniae ) and the goat moth (Cossus cossus) which have gained popularity as pests. However, the large caterpillars of species that do not smell badly are often edible. Witchetty grubs – among the Outback's most famous bush tucker – are most commonly the caterpillars of Endoxyla leucomochla , one of the more than 80 cossid species in Australia. In Chile, the sweet-smelling caterpillars of the Chilean moth ( Chilecomadia moorei ) are harvested in quantity and internationally traded as butterworms, for use as pet food and fishing bait.
Some other animal families, such as the Dudgeoneidae, Metarbelidae, and Ratardidae, have been included within this family time and again. The first is considered a distinct family of the Cossoidea today recognizable by their abdominal tympanal organs which the Cossidae lack, whereas the other two are usually kept in the Cossidae as subfamilies. Some unrelated millers were included in the Cossidae in error, too, such as the genus Holcoceroides which is more primitive Ditrysia, or the Andesianidae which are even more ancient Heteroneura.
The Cossidae were usually divided into six subfamilies. [2] However, numerous new subfamilies have been described recently, the current taxonomy is:
The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek geo γεω, and metron μέτρον "measure" in reference to the way their larvae, or inchworms, appear to measure the earth as they move along in a looping fashion. Geometridae is a very large family, containing around 23,000 described species; over 1400 species from six subfamilies are indigenous to North America alone. A well-known member is the peppered moth, Biston betularia, which has been the subject of numerous studies in population genetics. Several other geometer moths are notorious pests.
Aluctoidea is the superfamily of many-plumed and false plume moths. These small moths are most easily recognized by their wings. These each consist of many narrow strips of membrane around the major veins, instead of a continuous sheet of membrane between the veins. In living moths in the wild, this is often hard to see however. When they are at rest, the "plumes" partly overlap, appearing as solid wings. But even then, they can be recognized by the wings having a marked lengthwise pattern and uneven edge.
The Lasiocampidae are a family of moths also known as eggars, tent caterpillars, snout moths, or lappet moths. Over 2,000 species occur worldwide, and probably not all have been named or studied. It is the sole family in superfamily Lasiocampoidea.
The Coleophoridae are a family of small moths, belonging to the huge superfamily Gelechioidea. Collectively known as case-bearers, casebearing moths or case moths, this family is represented on all continents, but the majority are found in temperate areas of the Northern Hemisphere. They are most common in the Palearctic, and rare in sub-Saharan Africa, South America, and Australia; consequently, they probably originated in northern Eurasia. They are relatively common in houses, they seek out moist areas to rest and procreate.
The Dalceridae are a small family of zygaenoid moths with some 80 known species encompassing about one dozen genera mostly found in the Neotropical region with a few reaching the far south of the Nearctic region.
Xestia is a genus of noctuid moths. They are the type genus of the tribe Xestiini in subfamily Noctuinae, though some authors merge this tribe with the Noctuini. Species in this genus are commonly known as "clays", "darts" or "rustics", but such names are commonplace among Noctuidae. Xestia moths have a wide distribution, though they most prominently occur in the Holarctic.
The Ratardinae are a small subfamily of large moths from Southeast Asia.
The Cossinae are the nominate subfamily of the Cossidae. The caterpillars of several Cossinae species, such as the carpenterworm and the goat moth, are significant pests. On the other hand, in Chile the caterpillars of the Chilean moth are collected on a commercial scale for sale as fishing bait and terrarium pet food; they are usually called "butterworms" in international trade.
The Nacophorini are one of the smaller tribes of geometer moths in the subfamily Ennominae. They are the most diverse Ennominae of Australia and are widespread in the Americas. If the African genera tentatively placed herein indeed belong here, the distribution of the Nacophorini is distinctly Gondwanan, with their probable origin either of Australia, South America or even Antarctica. In Eurasia, they are rare by comparison.
Lomographa bimaculata, the white-pinion spotted, is a species of geometer moth. It belongs to the large geometer moth subfamily Ennominae, and therein to the tribe Baptini. It is – under its junior synonym – the type species of its genus Lomographa. It is also the type species of Bapta, a junior objective synonym of Lomographa and the namesake of the Baptini. The species was first described by Johan Christian Fabricius ion 1775.
Scythropia crataegella, the hawthorn moth, is a species of moth in the family Plutellidae from western Eurasia. It is usually placed in a small subfamily Scythropiinae, which is sometimes included in the Yponomeutinae of the Yponomeutidae.
Cossus cossus, the goat moth, is a moth of the family Cossidae. It is found in Northern Africa, Asia and Europe.
Oecophora bractella is a species of gelechioid moth. It belongs to the subfamily Oecophorinae of the concealer moth family (Oecophoridae). As the type species of its genus Oecophora, its affiliations and phylogeny determine the delimitation of that family and subfamily.
Borkhausenia minutella is a species of moth. Within its superfamily, it is placed within the subfamily Oecophorinae of the "concealer moth" family, Oecophoridae.
Alabonia geoffrella is a species of gelechioid moth. Here, it is placed within the subfamily Oecophorinae of the concealer moth family (Oecophoridae). Alternatively it has been placed in the Elachistidae or Depressariinae together with its presumed closest relatives.
Schiffermuelleria is a genus of gelechioid moths. It is placed in the subfamily Oecophorinae of family Oecophoridae. The genus is treated as monotypic, with the single species Schiffermuelleria schaefferella placed here. As such, its distinctness from the closely related genus Borkhausenia – where S. schaefferella was often placed in the past – is open to debate.
Scythris is a genus of gelechioid moths. It is the type genus of the flower moth family, which is sometimes included as a subfamily in the Xyloryctidae, or together with these merged into the Oecophoridae. The genus was erected by Jacob Hübner in 1825.
The Phycitinae are a subfamily of snout moths. Even though the Pyralidae subfamilies are all quite diverse, Phycitinae stand out even by standards of their family: with over 600 genera considered valid and more than 4000 species placed here at present, they unite up more than three-quarters of living snout moth diversity. Together with the closely related Epipaschiinae, they are apparently the most advanced lineage of snout moths.
The Galleriinae are a subfamily of snout moths and occur essentially worldwide, in some cases aided by involuntary introduction by humans. This subfamily includes the wax moths, whose caterpillars (waxworms) are bred on a commercial scale as food for pets and as fishing bait; in the wild, these and other species of Galleriinae may also be harmful to humans as pests.
Cossus is a genus of moths in the family Cossidae described by Johan Christian Fabricius in 1793.