180,000 species of Lepidoptera are described, equivalent to 10% of the total described species of living organisms. This is a list of the diversity of the Lepidoptera showing the estimated number of genera and species described for each superfamily and, where available, family. See Lepidoptera for a note of the schedule of families used.
Heterobathmia is a genus of Lepidoptera. It is the only genus in the suborder Heterobathmiina, as well as in the superfamily Heterobathmioidea and in the family Heterobathmiidae. Primitive, day-flying, metallic moths confined to southern South America, the adults eat the pollen of Nothofagus or southern beech and the larvae mine the leaves. Most known species are undescribed.
Simaethistoidea is an obscure superfamily of pyralid-like moths with two genera, whose biology and relationships among the Ditrysia is currently unknown, namely the Australian Metaprotus and the China and North Indian Simaethistis.
Neopseustidae is a small family of day and night-flying "archaic bell moths" in the order Lepidoptera. They are classified into their own superfamily Neopseustoidea and infraorder Neopseustina. Four genera are known. These primitive moths are restricted to South America and Southeast Asia. Their biology is unknown.
Immoidea is a superfamily of pantropical moths containing only the family Immidae comprising ten genera with around 250 species, over half of them in the genus Imma. Many are brightly coloured and diurnal. The position of this group is currently uncertain within the group Obtectomera. The larvae feed on the leaves of dicotyledons and conifers including Podocarpus.
Lophocoronoidea is a superfamily of insects in the order Lepidoptera. There is a single extant genus, Lophocorona, in the family Lophocoronidae. These are small, primitive nocturnal moths restricted to Australia whose biology is largely unknown.
Adeloidea is a superfamily of primitive monotrysian moths in the order Lepidoptera which consists of leafcutters, yucca moths and relatives. This superfamily is characterised by a piercing, extensible ovipositor used for laying eggs in plants. Many species are day-flying with metallic patterns.
Tischerioidea is the superfamily of "trumpet" leaf miner moths. The superfamily contains just one family, Tischeriidae, and traditionally one genus, Tischeria, but currently three genera are recognised, widespread around the world including South America, except for Australasia. This is one candidate as the sister group of the bulk of Lepidoptera, the Ditrysia, and they have a monotrysian type of female reproductive system. These small moths are leaf-miners in the caterpillar stage, feeding mainly on Fagaceae, Asteraceae, and Malvaceae (Astrotischeria), and some also on Rhamnaceae, Tiliaceae, and Rosaceae.
Callidulidae, the only known family of the superfamily Calliduloidea, is the family of Old World butterfly-moths, containing eight genera. They have a peculiar distribution, restricted to the Old World tropics of Southeast Asia to Australasia and Madagascar. The three subfamilies exhibit both day- and night-flying behaviour.
Carposinidae, the "fruitworm moths", is a family of insects in the order Lepidoptera. These moths are narrower winged than Copromorphidae, with less rounded forewing tips. Males often have conspicuous patches of scales on either surface. The mouthparts are quite diagnostic, usually with prominent, upcurved "labial palps", the third segment long, and the second segment covered in large scales. Unlike Copromorphidae, the "M2" and sometimes "M1" vein on the hindwings is absent. The relationship of Carposinidae relative to Copromorphidae needs further investigation. It is considered possible that the family is artificial, being nested within Copromorphidae. The Palearctic species have been revised by Alexey Diakonoff (1989).
Choreutidae, or metalmark moths, are a family of insects in the lepidopteran order whose relationships have been long disputed. It was placed previously in the superfamily Yponomeutoidea in family Glyphipterigidae and in superfamily Sesioidea. It is now considered to represent its own superfamily. The relationship of the family to the other lineages in the group "Apoditrysia" need a new assessment, especially with new molecular data.
Copromorphidae, the "tropical fruitworm moths", is a family of insects in the lepidopteran order. These moths have broad, rounded forewings, and well-camouflaged scale patterns. Unlike Carposinidae the mouthparts include "labial palps" with the second rather than third segment the longest. With other unusual structural characteristics of the caterpillar and adult, it could represent the sister lineage of all other extant members of this superfamily. The genus Sisyroxena from Madagascar is also notable for its unusual venation and wing scale sockets.
Hyblaeidae are the "teak moths", a family of insects in the Lepidopteran order. The two genera with about 18 species make up one of the two families of the Hyblaeoidea superfamily, which in the past has been included in the Pyraloidea. Recent phylogenetic studies find varying relationships of Hyblaeoidea among Ditrysian Lepidoptera: Mutanen et al. (2010) find the superfamily to group either with Pyraloidea, or – more often – with Thyridoidea or butterflies. The results of Wahlberg et al. (2013) and Heikilä et al. (2015) indicate a sister-group relationship with Pyraloidea.
Galacticidae is a recently recognised and enigmatic family of insects in the lepidopteran order. These moderate sized moths are 8–17 mm in wingspan and have previously been embedded within several lepidopteran superfamilies, but Galacticidae is currently placed in its own superfamily at the base of the natural group Apoditrysia.
Epermeniidae or the fringe-tufted moths is a family of insects in the lepidopteran order with about 14 genera. Previously they have been divided in two subfamilies Epermeniinae and Ochromolopinae but this is no longer maintained since the last group is probably hierarchically nested within the first. They are presently placed in their own superfamily but have previously been placed among the Yponomeutoidea or Copromorphoidea with which they share some features. Their systematic placement among the apoditrysian group "Obtectomera" is however uncertain. They show some morphological similarities to the "plume moths", for example the wing fringe has similar groups of scales. There are also some similarities to Schreckensteinioidea, for example spiny legs and at least in some species an open-network cocoon. The genus Thambotricha from New Zealand may be the sister group of all other extant members. The most important genera are Epermenia, Ochromolopis and Gnathifera. The group has been extensively revised and catalogued by Dr Reinhard Gaedike.
Whalleyana is an enigmatic genus of moths in the lepidopteran group Obtectomera, endemic to Madagascar. The genus contains two species, whose biology are unknown. The genus had been placed in the picture-winged leaf moths, (Thyrididae), but then was placed in its own family, and later elevated to its own superfamily ; see also Fänger (2004). The genus was named after Paul E. S. Whalley, a British entomologist. Genomic studies have found them to be most closely related to Callidulidae, and it is suggested that they should be placed in Calliduloidea.
Carposinoidea, the "fruitworm moths", is a superfamily of insects in the lepidopteran order. The superfamily is also known as Copromorphoidea, which is a junior synonym. These moths are small to medium-sized and are broad-winged bearing some resemblance to the superfamilies Tortricoidea and Immoidea. The antennae are often "pectinate" especially in males, and many species of these well camouflaged moths bear raised tufts of scales on the wings and a specialised fringe of scales at the base of the hindwing sometimes in females only; there are a number of other structural characteristics. The position of this superfamily is not certain, but it has been placed in the natural group of "Apoditrysia" "Obtectomera", rather than with the superfamilies Alucitoidea or Epermenioidea within which it has sometimes previously been placed, on the grounds that shared larval and pupal characteristics of these groups have probably evolved independently. It has been suggested that the division into two families should be abandoned.
The Palaeosetidae or miniature ghost moths are a family of insects in the order Lepidoptera contained within the superfamily Hepialoidea.
Acanthopteroctetidae is a small family of primitive moths with two described genera, Acanthopteroctetes and Catapterix, and a total of seven described species. They are known as the archaic sun moths.
Micropterigoidea is the superfamily of "mandibulate archaic moths", all placed in the single family Micropterigidae, containing currently about twenty living genera. They are considered the most primitive extant lineage of lepidoptera, and the sole superfamily in the suborder Zeugloptera. The name comes from the Greek for mikros, little and pterux, a wing. Unique among the Lepidoptera, these moths have chewing mouthparts rather than a proboscis, and are seen feeding, often in large aggregations, on the pollen of the flowers of many herbaceous plants, shrubs and trees. The fossil record of the group goes back to the middle-late Jurassic with the earliest known species being Auliepterix from the Karabastau Formation in Kazakhstan.
