| Sematuridae | |
|---|---|
 | |
| Mania empedocles | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Lepidoptera |
| Superfamily: | Geometroidea |
| Family: | Sematuridae |
| Subfamilies | |
| Diversity | |
| about 41 species | |
Sematuridae is a family of moths in the lepidopteran order that contains two subfamilies (Minet and Scoble, 1999).
These are large day- or night-flying moths, usually tailed and similar to Uraniidae in general appearance (except for the genera Apoprogones, Anuropteryx and Lonchotura). The position of this family is not certain amongst the Macrolepidoptera but it is usually considered to belong to the superfamily Geometroidea, whilst the identity of its closest extant relative is under investigation using DNA sequencing . Until recently very little has been known of sematurid biology (but see Sematurinae). The "chaetosemata" of Sematuridae have long setae overhanging the compound eyes, the antennae are thickened towards the often rather clubbed or hooked tip whilst a tympanal organ for hearing is always absent and there are other diagnostic characters in the abdomen (Minet and Scoble, 1999: 305-307; Holloway et al., 2001).
Apoprogoninae is represented by a single species in South Africa whilst Sematurinae is represented by about 29 (41 including the genera Anurapteryx and Lonchotura) species in the Neotropics. [1] Such an apparently relictual distribution might relate to the geological split of South America and Africa, but there is as yet no evidence for the age of evolutionary divergence of Neotropical and Afrotropical representatives; moreover, new genetic material would be needed for the South African taxon as well as the American genera Anurapteryx and Lonchotura for a modern molecular approach to this problem to succeed .
The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek geo γεω, and metron μέτρον "measure" in reference to the way their larvae, or inchworms, appear to measure the earth as they move along in a looping fashion. A very large family, it has around 23,000 species of moths described, and over 1400 species from six subfamilies indigenous to North America alone. A well-known member is the peppered moth, Biston betularia, which has been subject of numerous studies in population genetics. Several other geometer moths are notorious pests.
The Uraniidae are a family of moths containing four subfamilies, 90 genera, and roughly 700 species. The family is distributed throughout the tropics of the Americas, Africa and Indo-Australia. Some of the tropical species are known for their bright, butterfly-like colors and are called sunset moths. Such moths are apparently toxic and the bright colors are a warning to predators.
Riodinidae is the family of metalmark butterflies. The common name "metalmarks" refers to the small, metallic-looking spots commonly found on their wings. The 1,532 species are placed in 146 genera. Although mostly Neotropical in distribution, the family is also represented both in the Nearctic, Palearctic, Australasian (Dicallaneura), Afrotropic, and Indomalayan realms.
The Hepialidae are a family of insects in the lepidopteran order. Moths of this family are often referred to as swift moths or ghost moths.
The Drepanidae are a family of moths with about 660 species described worldwide. They are generally divided in three subfamilies, which share the same type of hearing organ. Thyatirinae, previously often placed in their own family, bear a superficial resemblance to Noctuidae. Many species in the drepanid family have a distinctively hook-shaped apex to the fore wing, leading to their common name of hook-tips.
Schreckensteinioidea is a superfamily in the insect order Lepidoptera containing a single family, Schreckensteiniidae, or "bristle-legged moths", because of the stout spines on the hindlegs. The superfamily and family were both described by Thomas Bainbrigge Fletcher in 1929. The relationships of this family within the group apoditrysia are currently uncertain. One of the species, the blackberry skeletoniser, is widespread and common across Europe and has been introduced as a biological control to Hawaii, whilst three species of Corsocasis occur in South East Asia.
Nepticulidae is a family of very small moths with a worldwide distribution. They are characterised by eyecaps over the eyes. These pigmy moths or midget moths, as they are commonly known, include the smallest of all living moths, with a wingspan that can be as little as 3 mm in the case of the European pigmy sorrel moth, but more usually 3.5–10 mm. The wings of adult moths are narrow and lanceolate, sometimes with metallic markings, and with the venation very simplified compared to most other moths.
Callidulidae, the only known family of the superfamily Calliduloidea, is the family of Old World butterfly-moths, containing eight genera. They have a peculiar distribution, restricted to the Old World tropics of Southeast Asia to Australasia and Madagascar. The three subfamilies exhibit both day- and night-flying behaviour.
Carposinidae, the "fruitworm moths", is a family of insects in the order Lepidoptera. These moths are narrower winged than Copromorphidae, with less rounded forewing tips. Males often have conspicuous patches of scales on either surface. The mouthparts are quite diagnostic, usually with prominent, upcurved "labial palps", the third segment long, and the second segment covered in large scales. Unlike Copromorphidae, the "M2" and sometimes "M1" vein on the hindwings is absent. The relationship of Carposinidae relative to Copromorphidae needs further investigation. It is considered possible that the family is artificial, being nested within Copromorphidae. The Palearctic species have been revised by Alexey Diakonoff (1989).
Choreutidae, or metalmark moths, are a family of insects in the lepidopteran order whose relationships have been long disputed. It was placed previously in the superfamily Yponomeutoidea in family Glyphipterigidae and in superfamily Sesioidea. It is now considered to represent its own superfamily. The relationship of the family to the other lineages in the group "Apoditrysia" need a new assessment, especially with new molecular data.
Copromorphidae, the "tropical fruitworm moths", is a family of insects in the lepidopteran order. These moths have broad, rounded forewings, and well-camouflaged scale patterns. Unlike Carposinidae the mouthparts include "labial palps" with the second rather than third segment the longest. With other unusual structural characteristics of the caterpillar and adult, it could represent the sister lineage of all other extant members of this superfamily. The genus Sisyroxena from Madagascar is also notable for its unusual venation and wing scale sockets.
Hyblaeidae are the "teak moths", a family of insects in the Lepidopteran order. The two genera with about 18 species make up one of the two families of the Hyblaeoidea superfamily, which in the past has been included in the Pyraloidea. Recent phylogenetic studies find varying relationships of Hyblaeoidea among Ditrysian Lepidoptera: Mutanen et al. (2010) find the superfamily to group either with Pyraloidea, or – more often – with Thyridoidea or butterflies. The results of Wahlberg et al. (2013) and Heikilä et al. (2015) indicate a sister-group relationship with Pyraloidea.
Hedylidae, the "American moth-butterflies", is a family of insects in the order Lepidoptera, representing the superfamily Hedyloidea. They have traditionally been viewed as an extant sister group of the butterfly superfamily Papilionoidea, but a 2014 phylogenetic analyses has suggested Hedylidae is a subgroup of Papilionoidea, and not a sister group, and are more accurately referred to as butterflies rather than moths. They are represented by a single Neotropical genus Macrosoma with 35 currently recognized species.
Galacticidae is a recently recognised and enigmatic family of insects in the lepidopteran order. These moderate sized moths are 8–17 mm in wingspan and have previously been embedded within several lepidopteran superfamilies, but Galacticidae is currently placed in its own superfamily at the base of the natural group Apoditrysia.
Epermeniidae or the fringe-tufted moths is a family of insects in the lepidopteran order with about 14 genera. Previously they have been divided in two subfamilies Epermeniinae and Ochromolopinae but this is no longer maintained since the last group is probably hierarchically nested within the first. They are presently placed in their own superfamily but have previously been placed among the Yponomeutoidea or Copromorphoidea with which they share some features. Their systematic placement among the apoditrysian group "Obtectomera" is however uncertain. They show some morphological similarities to the "plume moths", for example the wing fringe has similar groups of scales. There are also some similarities to Schreckensteinioidea, for example spiny legs and at least in some species an open-network cocoon. The genus Thambotricha from New Zealand may be the sister group of all other extant members. The most important genera are Epermenia, Ochromolopis and Gnathifera. The group has been extensively revised and catalogued by Dr Reinhard Gaedike.
Whalleyana is an enigmatic genus of moths in the lepidopteran group Obtectomera, endemic to Madagascar. The genus contains two species, whose biology are unknown. The genus had been placed in the picture-winged leaf moths, (Thyrididae), but then was placed in its own family, and later elevated to its own superfamily ; see also Fänger (2004). The genus was named after Paul E. S. Whalley, a British entomologist. Genomic studies have found them to be most closely related to Callidulidae, and it is suggested that they should be placed in Calliduloidea.
Carposinoidea, the "fruitworm moths", is a superfamily of insects in the lepidopteran order. The superfamily is also known as Copromorphoidea, which is a junior synonym. These moths are small to medium-sized and are broad-winged bearing some resemblance to the superfamilies Tortricoidea and Immoidea. The antennae are often "pectinate" especially in males, and many species of these well camouflaged moths bear raised tufts of scales on the wings and a specialised fringe of scales at the base of the hindwing sometimes in females only; there are a number of other structural characteristics. The position of this superfamily is not certain, but it has been placed in the natural group of "Apoditrysia" "Obtectomera", rather than with the superfamilies Alucitoidea or Epermenioidea within which it has sometimes previously been placed, on the grounds that shared larval and pupal characteristics of these groups have probably evolved independently. It has been suggested that the division into two families should be abandoned.
Pterolonchidae is a small family of very small moths in the superfamily Gelechioidea. There are species native to every continent except Australia and Antarctica.
Apoprogoninae is a monotypic subfamily of the moth family Sematuridae. Its single genus, Apoprogones, containing a single species, Apoprogones hesperistis, were both described by George Hampson in 1903. It is known from Eswatini and South Africa.
Sterrhinae is a large subfamily of geometer moths with some 3,000 described species, with more than half belonging to the taxonomically difficult, very diverse genera, Idaea and Scopula. This subfamily was described by Edward Meyrick in 1892. They are the most diverse in the tropics with the number of species decreasing with increasing latitude and elevation.
