Taxonomy
Around the start of the century, they were considered the fifth group up on the comb of branching events in the extant lepidopteran phylogeny, [9] : 10 and also deemed to represent the most basal lineage in the lepidopteran group Coelolepida [10] (along with Lophocoronoidea and the massive group "Myoglossata") characterised in part by its scale morphology. [3] : 53–54
Research on the molecular phylogeny of the Lepidoptera since then has indicated a close relation between the Acanthopteroctetidae, the Neopseustidae and the Aenigmatineidae, [1] : 672–681 and the three may be considered part of a single superfamily Neopseustoidea rather than three separate, monobasic superfamilies. [1] : 681 Molecular data from the same research showed weak support for the clade Coelolepida, and weakly contradicted the placement of Acanthopteroctetidae as most basal lineage of the Coelolepida. [1] : 676
Genus Catapterix was originally described within its own family, Catapterigidae, [11] [12] which is considered a junior synonym of Acanthopteroctidae, [13] with which it shares specialised structural features including similar wing morphology (in A. unifascia). [4] : 1255
The Ditrysia are a natural group or clade of insects in the lepidopteran order containing both butterflies and moths. They are so named because the female has two distinct sexual openings: one for mating, and the other for laying eggs.
Neopseustidae is a small family of day and night-flying "archaic bell moths" in the order Lepidoptera. They are classified into their own superfamily Neopseustoidea and infraorder Neopseustina. Four genera are known. These primitive moths are restricted to South America and Southeast Asia. Their biology is unknown.
Eriocraniidae is a family of moths restricted to the Holarctic region, with six extant genera. These small, metallic moths are usually day-flying, emerging fairly early in the northern temperate spring. They have a proboscis with which they drink water or sap. The larvae are leaf miners on Fagales, principally the trees birch (Betula) and oak (Quercus), but a few on Salicales and Rosales.
Lophocoronoidea is a superfamily of insects in the order Lepidoptera. There is a single extant genus, Lophocorona, in the family Lophocoronidae. These are small, primitive nocturnal moths restricted to Australia whose biology is largely unknown.
The Hepialoidea are the superfamily of "ghost moths" and "swift moths".
Palaephatoidea is a superfamily of insects in the order Lepidoptera with a single family, Palaephatidae with seven known genera. These "Gondwanaland moths" exhibit a disjunct distribution occurring mainly in South America, with four species in eastern Australia and Tasmania and one in South Africa. The larvae spin together leaves of Proteaceae (Ptyssoptera) or Verbenaceae (Azaleodes) . Palaephatoidea, a typical monotrysian group, is one two main candidates as the sister group of most of the Lepidoptera, the Ditrysia.
Tischerioidea is the superfamily of "trumpet" leaf miner moths. The superfamily contains just one family, Tischeriidae, and traditionally one genus, Tischeria, but currently three genera are recognised, widespread around the world including South America, except for Australasia. This is one candidate as the sister group of the bulk of Lepidoptera, the Ditrysia, and they have a monotrysian type of female reproductive system. These small moths are leaf-miners in the caterpillar stage, feeding mainly on Fagaceae, Asteraceae, and Malvaceae (Astrotischeria), and some also on Rhamnaceae, Tiliaceae, and Rosaceae.
Coelolepida is a clade of insects in the lepidopteran order, containing the infraorders Acanthoctesia and Lophocoronina.
The Adelidae or fairy longhorn moths are a family of monotrysian moths in the lepidopteran infraorder Heteroneura. The family was first described by Charles Théophile Bruand d'Uzelle in 1851. Most species have at least partially metallic, patterned coloration and are diurnal, sometimes swarming around the tips of branches with an undulating flight. Others are crepuscular and have a drab coloration. Fairy longhorn moths have a wingspan of 4–28 millimeters, and males often have especially long antennae, 1–3 times as long as the forewing.
Hyblaeidae are the "teak moths", a family of insects in the Lepidopteran order. The two genera with about 18 species make up one of the two families of the Hyblaeoidea superfamily, which in the past has been included in the Pyraloidea. Recent phylogenetic studies find varying relationships of Hyblaeoidea among Ditrysian Lepidoptera: Mutanen et al. (2010) find the superfamily to group either with Pyraloidea, or – more often – with Thyridoidea or butterflies. The results of Wahlberg et al. (2013) and Heikilä et al. (2015) indicate a sister-group relationship with Pyraloidea.
Whalleyana is an enigmatic genus of moths in the lepidopteran group Obtectomera, endemic to Madagascar. The genus contains two species, whose biology are unknown. The genus had been placed in the picture-winged leaf moths, (Thyrididae), but then was placed in its own family, and later elevated to its own superfamily ; see also Fänger (2004). The genus was named after Paul E. S. Whalley, a British entomologist. Genomic studies have found them to be most closely related to Callidulidae, and it is suggested that they should be placed in Calliduloidea.
Carposinoidea, the "fruitworm moths", is a superfamily of insects in the lepidopteran order. The superfamily is also known as Copromorphoidea, which is a junior synonym. These moths are small to medium-sized and are broad-winged bearing some resemblance to the superfamilies Tortricoidea and Immoidea. The antennae are often "pectinate" especially in males, and many species of these well camouflaged moths bear raised tufts of scales on the wings and a specialised fringe of scales at the base of the hindwing sometimes in females only; there are a number of other structural characteristics. The position of this superfamily is not certain, but it has been placed in the natural group of "Apoditrysia" "Obtectomera", rather than with the superfamilies Alucitoidea or Epermenioidea within which it has sometimes previously been placed, on the grounds that shared larval and pupal characteristics of these groups have probably evolved independently. It has been suggested that the division into two families should be abandoned.
Catapterix is a small genus of moths in the family Acanthopteroctetidae, with species occurring in Europe and Central Asia. The genus was previously considered monotypic, with Catapterix crimaea as its sole known species, until the formal description of Catapterix tianshanica in 2016.
Heteroneura is a natural group in the insect order Lepidoptera that comprises over 99% of all butterflies and moths. This is the sister group of the infraorder Exoporia, and is characterised by wing venation which is not similar or homoneurous in both pairs of wings. Though basal groups within the Heteroneura cannot be identified with much confidence, one major subgroup is the leaf-mining Nepticuloidea. Species in this subgroup include some of the smallest lepidopterans identified.
The insect order Lepidoptera consists of moths, most of which are night-flying, and a derived group, mainly day-flying, called butterflies. Within Lepidoptera as a whole, the groups listed below before Glossata contain a few basal families accounting for less than 200 species; the bulk of Lepidoptera are in the Glossata. Similarly, within the Glossata, there are a few basal groups listed first, with the bulk of species in the Heteroneura. Basal groups within Heteroneura cannot be defined with as much confidence, as there are still some disputes concerning the proper relations among these groups. At the family level, however, most groups are well defined, and the families are commonly used by hobbyists and scientists alike.
Catapterix crimaea is a moth of the family Acanthopteroctetidae and type species of the genus Catapterix. It was described in 1988 from the Crimean peninsula of Ukraine, and was in 2016 first recorded from southern France. From its description in 1988 up until the description of Catapterix tianshanica in 2016, Catapterix crimaea was considered the sole species of its genus.
Acanthopteroctetes bimaculata is a moth of the family Acanthopteroctetidae. It was described by Davis in 1969. It is found in north-eastern Oregon and east-central California.
Acanthopteroctetes unifascia is a moth of the family Acanthopteroctetidae. It was described by Davis in 1978. It is found in Montana.
Tridentaforma is a genus of moths of the Tridentaformidae family, the sole genus of the family. There is only one species in the genus, Tridentaforma fuscoleuca, which is found in California and Canada.
Aenigmatineidae is a family of basal Lepidoptera, moths discovered on Kangaroo Island in South Australia by Dr Richard Glatz. The family is based on a single species discovered in 2015, Aenigmatinea glatzella, commonly known as the enigma moth. The larvae feed on conifers by mining the stem of Callitris plants in the cypress family. The adult has highly reduced mouthparts but its position in the Glossata containing the more familiar moths-with-tongues is confirmed by morphological and DNA sequence similarity. The group is best treated as a sister of the family Neopseustidae.
