Callidulidae, the only known family of the superfamilyCalliduloidea, is the family of Old World butterfly-moths, containing eight genera. They have a peculiar distribution, restricted to the Old World tropics of Southeast Asia to Australasia and Madagascar. The three subfamilies exhibit both day- and night-flying behaviour.
The mainly day-flying Callidulinae can be distinguished by their resting posture, which is the most butterfly-like, with the wings held closely over the back. Resembling the butterfly familyLycaenidae, these moths can be told apart by their antennae which taper to a point or may be very subtly clubbed. The more often night-flying Pterothysaninae and Griveaudiinae have a different adult resting posture (the latter roof-like in repose) and these were not placed within the Callidulidae until recently.[1]
Biology of most subfamilies and species is poorly known. Eggs are very flat in Griveaudiinae and Callidulinae, and caterpillars of Callidulinae are green with a shiny black head and have only been reported from ferns[2] forming a leaf roll in which they live, eat and finally pupate, while the pupa of Helicomitra appears to be subterranean.[3]
The closest relatives of Callidulidae are not known, but they are currently placed in a group that includes the three butterfly superfamilies, the "hook-tip moths" Drepanoidea and the "geometer moths" Geometroidea and also possibly Axioidea which share some structural characteristics.[4]
Cited references
↑ Minet, J. (1986). Ébauche d'une classification moderne de l'ordre des Lépidoptères. Alexanor 14(7): 291–313.
↑ Gaden S. Robinson, Phillip R. Ackery, Ian J. Kitching, George W. Beccaloni & Luis M. Hernández. HOSTS database. (Accessed May 2007)
↑ Minet, J. (1987). Description d'une chrysalide de Pterothysaninae (Lep. Callidulidae). Nouvelle Revue d'Entomologie (N.S.) 4(3): 312.
↑ Minet, J. (1999 [1988]). The Axioidea and Calliduloidea. Ch. 16, pp. 257–261 in Kristensen, N.P. (Ed.). Lepidoptera, Moths and Butterflies. Volume 1: Evolution, Systematics, and Biogeography. Handbuch der Zoologie. Eine Naturgeschichte der Stämme des Tierreiches / Handbook of Zoology. A Natural History of the phyla of the Animal Kingdom. Band / Volume IV Arthropoda: Insecta Teilband / Part 35: 491 pp. Walter de Gruyter, Berlin, New York.
Other references
Minet, J. (1990 [1989]). Nouvelles frontières, géographiques et taxonomiques, pour la famille des Callidulidae (Lepidoptera, Calliduloidea). Nouvelle Revue d'Entomologie (N.S.) 6(4): 351–368.
Maddison, David R. 2003. Callidulidae. in The Tree of Life Web Project, Accessed October 2006
O'Toole, Christopher (Ed.) (2002) Firefly Encyclopedia of Insects and Spiders. ISBN1-55297-612-2
Crambidae comprises the grass moth family of lepidopterans. They are variable in appearance, with the nominal subfamily Crambinae taking up closely folded postures on grass stems where they are inconspicuous, while other subfamilies include brightly coloured and patterned insects that rest in wing-spread attitudes.
Macrolepidoptera is a group within the insect order Lepidoptera. Traditionally used for the larger butterflies and moths as opposed to the "microlepidoptera", this group is artificial. However, it seems that by moving some taxa about, a monophyletic macrolepidoptera can be easily achieved. The two superfamilies Geometroidea and Noctuoidea account for roughly one-quarter of all known Lepidoptera.
The Drepanidae are a family of moths with about 660 species described worldwide. They are generally divided in three subfamilies, which share the same type of hearing organ. Thyatirinae, previously often placed in their own family, bear a superficial resemblance to Noctuidae. Many species in the drepanid family have a distinctively hook-shaped apex to the fore wing, leading to their common name of hook-tips.
Neopseustidae is a small family of day and night-flying "archaic bell moths" in the order Lepidoptera. They are classified into their own superfamily Neopseustoidea and infraorder Neopseustina. Four genera are known. These primitive moths are restricted to South America and Southeast Asia. Their biology is unknown.
Eriocraniidae is a family of moths restricted to the Holarctic region, with six extant genera. These small, metallic moths are usually day-flying, emerging fairly early in the northern temperate spring. They have a proboscis with which they drink water or sap. The larvae are leaf miners on Fagales, principally the trees birch (Betula) and oak (Quercus), but a few on Salicales and Rosales.
Gelechioidea is the superfamily of moths that contains the case-bearers, twirler moths, and relatives, also simply called curved-horn moths or gelechioid moths. It is a large and poorly understood '"micromoth" superfamily, constituting one of the basal lineages of the Ditrysia.
Aluctoidea is the superfamily of many-plumed and false plume moths. These small moths are most easily recognized by their wings. These each consist of many narrow strips of membrane around the major veins, instead of a continuous sheet of membrane between the veins. In living moths in the wild, this is often hard to see however. When they are at rest, the "plumes" partly overlap, appearing as solid wings. But even then, they can be recognized by the wings having a marked lengthwise pattern and uneven edge.
Carposinidae, the "fruitworm moths", is a family of insects in the order Lepidoptera. These moths are narrower winged than Copromorphidae, with less rounded forewing tips. Males often have conspicuous patches of scales on either surface. The mouthparts are quite diagnostic, usually with prominent, upcurved "labial palps", the third segment long, and the second segment covered in large scales. Unlike Copromorphidae, the "M2" and sometimes "M1" vein on the hindwings is absent. The relationship of Carposinidae relative to Copromorphidae needs further investigation. It is considered possible that the family is artificial, being nested within Copromorphidae. The Palearctic species have been revised by Alexey Diakonoff (1989).
Choreutidae, or metalmark moths, are a family of insects in the lepidopteran order whose relationships have been long disputed. It was placed previously in the superfamily Yponomeutoidea in family Glyphipterigidae and in superfamily Sesioidea. It is now considered to represent its own superfamily. The relationship of the family to the other lineages in the group "Apoditrysia" need a new assessment, especially with new molecular data.
Hedylidae, the "American moth-butterflies", is a family of insects in the order Lepidoptera, representing the superfamily Hedyloidea. They have traditionally been viewed as an extant sister group of the butterfly superfamily Papilionoidea, but a 2014 phylogenetic analyses has suggested Hedylidae is a subgroup of Papilionoidea, and not a sister group, and are more accurately referred to as butterflies rather than moths. They are represented by a single Neotropical genus Macrosoma with 35 currently recognized species.
Galacticidae is a recently recognised and enigmatic family of insects in the lepidopteran order. These moderate sized moths are 8–17 mm in wingspan and have previously been embedded within several lepidopteran superfamilies, but Galacticidae is currently placed in its own superfamily at the base of the natural group Apoditrysia.
Epermeniidae or the fringe-tufted moths is a family of insects in the lepidopteran order with about 14 genera. Previously they have been divided in two subfamilies Epermeniinae and Ochromolopinae but this is no longer maintained since the last group is probably hierarchically nested within the first. They are presently placed in their own superfamily but have previously been placed among the Yponomeutoidea or Copromorphoidea with which they share some features. Their systematic placement among the apoditrysian group "Obtectomera" is however uncertain. They show some morphological similarities to the "plume moths", for example the wing fringe has similar groups of scales. There are also some similarities to Schreckensteinioidea, for example spiny legs and at least in some species an open-network cocoon. The genus Thambotricha from New Zealand may be the sister group of all other extant members. The most important genera are Epermenia, Ochromolopis and Gnathifera. The group has been extensively revised and catalogued by Dr Reinhard Gaedike.
Brachodidae is a family of day-flying moths, commonly known as little bear moths, which contains about 135 species distributed around much of the world. The relationships and status of the presently included genera are not well understood.
Whalleyana is an enigmatic genus of moths in the lepidopteran group Obtectomera, endemic to Madagascar. The genus contains two species, whose biology are unknown. The genus had been placed in the picture-winged leaf moths, (Thyrididae), but then was placed in its own family, and later elevated to its own superfamily ; see also Fänger (2004). The genus was named after Paul E. S. Whalley, a British entomologist. Genomic studies have found them to be most closely related to Callidulidae, and it is suggested that they should be placed in Calliduloidea.
Carposinoidea, the "fruitworm moths", is a superfamily of insects in the lepidopteran order. The superfamily is also known as Copromorphoidea, which is a junior synonym. These moths are small to medium-sized and are broad-winged bearing some resemblance to the superfamilies Tortricoidea and Immoidea. The antennae are often "pectinate" especially in males, and many species of these well camouflaged moths bear raised tufts of scales on the wings and a specialised fringe of scales at the base of the hindwing sometimes in females only; there are a number of other structural characteristics. The position of this superfamily is not certain, but it has been placed in the natural group of "Apoditrysia" "Obtectomera", rather than with the superfamilies Alucitoidea or Epermenioidea within which it has sometimes previously been placed, on the grounds that shared larval and pupal characteristics of these groups have probably evolved independently. It has been suggested that the division into two families should be abandoned.
Sematuridae is a family of moths in the lepidopteran order that contains two subfamilies.
The Ratardinae are a small subfamily of large moths from Southeast Asia.
The insect order Lepidoptera consists of moths, most of which are night-flying, and a derived group, mainly day-flying, called butterflies. Within Lepidoptera as a whole, the groups listed below before Glossata contain a few basal families accounting for less than 200 species; the bulk of Lepidoptera are in the Glossata. Similarly, within the Glossata, there are a few basal groups listed first, with the bulk of species in the Heteroneura. Basal groups within Heteroneura cannot be defined with as much confidence, as there are still some disputes concerning the proper relations among these groups. At the family level, however, most groups are well defined, and the families are commonly used by hobbyists and scientists alike.
Cimeliidae, the gold moths, is a family of moths that is now placed in the macroheteroceran superfamily Drepanoidea, although previously placed in its own superfamily. Uniquely, they have a pair of pocket-like organs on the seventh abdominal spiracle of the adult moth which are only possibly sound receptive organs. They are quite large and brightly coloured moths that occur in southern Europe and feed on species of Euphorbia. Sometimes they are attracted to light. The family was first described by Pierre Chrétien in 1916.
Scopariinae is a subfamily of the lepidopteran family Crambidae. The subfamily was described by Achille Guenée in 1854.
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