Guanay cormorant | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Suliformes |
Family: | Phalacrocoracidae |
Genus: | Leucocarbo |
Species: | L. bougainvilliorum |
Binomial name | |
Leucocarbo bougainvilliorum (Lesson, RP, 1837) | |
Distribution map of the guanay cormorant | |
Synonyms | |
Phalacrocorax bougainvillii |
The guanay cormorant or guanay shag (Leucocarbo bougainvilliorum) is a member of the cormorant family found on the Pacific coast of Peru and northern Chile. (The Argentinian population on the Patagonian Atlantic coast appears to be extirpated.) After breeding it spreads south to southern parts of Chile and north to Ecuador, and has also been recorded as far north as Panama and Colombia – probably a result of mass dispersal due to food shortage in El Niño years. Its major habitats include shallow seawater and rocky shores.
The guanay cormorant is similar in coloration to the rock shag, Leucocarbo magellanicus, but larger, measuring 78 cm from the tip of the bill to the end of the tail. Its bill is grayish with some red at the base. The face is red with a green eye-ring. It has roseate feet. The head, neck and back are black as are the outer parts of the thighs. The throat patch, breast and belly are white. In breeding plumage it has a few white feathers on the sides of the head and neck.
Breeding occurs year round with a peak in November and December. The nest is built of guano on flat surfaces on offshore islands or remote headlands. There are up to three nests per square meter in high-density colonies. The guanay cormorant lays two or three eggs of approximately 63 × 40 mm in size.
It feeds mainly on the Peruvian anchoveta, Engraulis ringens, and the Peruvian silverside, Odontesthes regia, which thrive in the cold Humboldt Current.[ citation needed ] The guanay cormorant is the main producer of guano. [2]
Habitat loss and degradation and over-fishing have resulted in a steady decline of the population of about 30% from an estimated figure of three million birds in 1984. This species is listed as near threatened by IUCN.
Some taxonomic authorities, including the International Ornithologists' Union, place this species in the genus Leucocarbo . Others place it in the genus Phalacrocorax .
The scientific name commemorates the French explorer Louis Antoine de Bougainville. The bird's droppings were such an important source of fertilizer to the peoples of the Andes that it was protected by Inca rulers, who supposedly made disturbing the cormorants in any way punishable by death. The common name is an adaptation of the South American Spanish guanae, a plural of the English equivalent guano. [3]
Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the International Ornithologists' Union (IOU) adopted a consensus taxonomy of seven genera. The great cormorant and the common shag are the only two species of the family commonly encountered in Britain and Ireland and "cormorant" and "shag" appellations have been later assigned to different species in the family somewhat haphazardly.
The pelagic cormorant, also known as Baird's cormorant or violet-green cormorant, is a small member of the cormorant family Phalacrocoracidae. Analogous to other smallish cormorants, it is also called the pelagic shag occasionally. This seabird lives along the coasts of the northern Pacific; during winter it can also be found in the open ocean. Pelagic cormorants have relatively short wings due to their need for economical movement underwater, and consequently have the highest flight costs of any bird.
The European shag or common shag is a species of cormorant. It is the only member of the monotypic genus Gulosus. It breeds around the rocky coasts of western and southern Europe, southwest Asia and north Africa, mainly wintering in its breeding range except for the northernmost birds. In Britain this seabird is usually referred to as simply the shag. The scientific genus name derives from the Latin for glutton. The species name aristotelis commemorates the Greek philosopher Aristotle.
The Australian pied cormorant, also known as the pied cormorant, pied shag, or great pied cormorant, is a medium-sized member of the cormorant family. It is found around the coasts of Australasia. In New Zealand, it is usually known either as the pied shag or by its Māori name of kāruhiruhi. Older sources may refer to it as the "yellow-faced cormorant".
The Otago shag, together with the Foveaux shag formerly known as the Stewart Island shag and in its dark phase as the bronze shag, is a species of shag now found only in coastal Otago, New Zealand.
The Cape cormorant or Cape shag is a bird endemic to the southwestern coasts of Africa.
The Socotra cormorant is a threatened species of cormorant that is endemic to the Persian Gulf and the south-east coast of the Arabian Peninsula. It is also sometimes known as the Socotran cormorant or, more rarely, as the Socotra shag. Individuals occasionally migrate as far west as the Red Sea coast. Despite its name, it was only confirmed in 2005 that it breeds on the Socotra islands in the Indian Ocean.
The rock shag, also known as the Magellanic cormorant, is a marine cormorant found around the southernmost coasts of South America. Its breeding range is from around Valdivia, Chile, south to Cape Horn and Tierra del Fuego, and north to Punta Tombo in Argentina. In winter it is seen further north, with individuals reaching as far as Santiago, Chile on the west coast and Uruguay on the east. The birds also breed around the coasts of the Falkland Islands.
The New Zealand king shag, also known as the rough-faced shag, king shag or kawau tūī, is a rare bird endemic to New Zealand. Some taxonomic authorities, including the International Ornithologists' Union, place this species in the genus Leucocarbo. Others place it in the genus Phalacrocorax.
The black-faced cormorant, also known as the black-faced shag, is a medium-sized member of the cormorant family. Upperparts, including facial skin and bill, are black, with white underparts. It is endemic to coastal regions of southern Australia.
The Auckland shag or Auckland Islands shag is a species of cormorant from New Zealand. The species is endemic to the Auckland Islands archipelago. It is a sedentary bird that primarily eats various crustaceans and fish. In recent years, roughly 1,000 pairs have been recorded. The Auckland shag is a colonial nester, building sizeable nests of, among other items, grass, twigs and seaweed. The Auckland shag lays three pale blue-green eggs in November–February. The incubation period is 26–32 days.
The imperial shag or imperial cormorant is a black and white cormorant native to southern South America, primarily in rocky coastal regions, but locally also at large inland lakes. Some taxonomic authorities, including the International Ornithologists' Union, place it in the genus Leucocarbo, others in the genus Phalacrocorax. It is also known as the blue-eyed shag, blue-eyed cormorant and by many other names, and is one of a larger group of cormorants called blue-eyed shags. The taxonomy is very complex, and several former subspecies are often considered separate species.
The Pitt shag, also known as the Pitt Island shag or Featherstone's shag, is a species of bird in the family Phalacrocoracidae. It is endemic to Pitt Island where its natural habitats are open seas and rocky shores. The bird with the color of a dark body, gray chest, yellow eyes and feet can be found in small groups in their breeding habitat or alone within 18 km from their territory searching for food. The species was close to extinction in 1905 but actions of conservation have been done ever since. The Pitt shag is threatened by habitat loss due to predation and climate change causing a decline from 1997 to 2012, but there is a recovery plan by the Department of Conservation.
The red-legged cormorant, also known as the red-legged shag, red-footed cormorant, red-footed shag, Gaimard's cormorant and grey cormorant, is a species of cormorant resident to the coastline of South America. It is the only member of the genus Poikilocarbo. It is non-colonial unlike most seabirds. The red-legged cormorant has not been observed wing-spreading, which is unusual among cormorant species.
The Chatham shag, also known as the Chatham Island shag, is a species of bird in the cormorant and shag family, Phalacrocoracidae. It is endemic to the Chatham Islands of New Zealand. For a long time the species was placed in the genus Phalacrocorax; today it is mostly placed with the other blue-eyed shags of New Zealand and Antarctica in the genus Leucocarbo. Its closest relative is the Otago shag of South Island.
The Bounty shag, also known as the Bounty Island shag, is a species of cormorant of the family Phalacrocoracidae. They are found only on the tiny and remote Subantarctic Bounty Islands, 670 km (420 mi) southeast of New Zealand. Its natural habitats are open seas and rocky shores. In 2022, a full Unmanned aerial vehicle survey of the Bounty archipelago found a total of 573 breeding pairs and estimates the population to consist of approximately 1,733 birds. These recent estimates are consistent with the only other comparable study from 1978 and suggest that the species' population has remained stable over the past 45 years.
The Kerguelen shag is a species of cormorant endemic to the Kerguelen Islands in the southern Indian Ocean, one of the most isolated places on Earth. Many authorities consider it a subspecies of the imperial shag.
The Macquarie shag, Macquarie Island shag or Macquarie Island cormorant, is a marine cormorant native to Macquarie Island in the Southern Ocean, about halfway between Australia and Antarctica.
The order Suliformes is an order of birds recognised by the International Ornithologist's Union. Regarding the recent evidence that the traditional Pelecaniformes is polyphyletic, it has been suggested that the group be divided to reflect the true evolutionary relationships; a 2017 study indicated that they are most closely related to Otidiformes (bustards) and Ciconiiformes (storks).
The Crozet shag, also known as the South Georgia cormorant, is a marine cormorant native to the Crozet, Prince Edward and Marion islands in the South Atlantic Ocean.