Black-faced cormorant | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Suliformes |
Family: | Phalacrocoracidae |
Genus: | Phalacrocorax |
Species: | P. fuscescens |
Binomial name | |
Phalacrocorax fuscescens (Vieillot, 1817) | |
Distribution of the black-faced cormorant | |
Synonyms | |
Stictocarbo fuscescens |
The black-faced cormorant (Phalacrocorax fuscescens), also known as the black-faced shag, is a medium-sized member of the cormorant family. Upperparts, including facial skin and bill, are black, with white underparts. It is endemic to coastal regions of southern Australia.
Like other cormorant species, the black-faced cormorant is a large aquatic bird, with a long hooked bill, webbed feet, and monochromatic plumage. [2] [3] This is one of the largest cormorants found in south-western Australia [2] and has pied plumage with the upper half of its body black and the undersides white. Its face is naked and black, hence the "black-faced" name, and the tail, feet, and thighs are also black. [4] The back feathers are glossy, and its bill is dark grey with a prominent hook at the tip. It has blue-green eyes. When flying, it holds its head level or lower than its body and holds its wings in a cross-shape like most cormorants. [4]
Species that are similar in appearance include the pied cormorant (slightly larger) and the little pied cormorant (slightly smaller). [5]
The breeding and non-breeding plumages are very similar, with the formation of fine white streaks along the neck during breeding until eggs are laid. [6] Juveniles have dark-brown upper feathers, including dark ear coverts and a dark face, with a lighter underside streaked with brown. Following the Juvenile plumage, black-faced cormorants have immature plumage which are similar to adult plumage. [4]
Black-faced cormorants have very subtle sexual dimorphism, making it difficult to differentiate sexes in the field. In general, males are larger and more robust than females, who are more slender in comparison. [7] Males also have larger bills than females, with male bills measuring greater than 77 millimeters and female bills less than 76 mm. [7]
Black-faced cormorants are usually silent when away from the breeding colony. [4] [7] [6] However, when they are at their nests, the male will emit a loud honk or guttural croak, and the female will hiss when approached. [7] [4]
The black-faced cormorant is one of around 40 species in the cormorant and shag family Phalacrocoracidae. [3] This family split off from the darter family Anhingadae over 40 million years ago, so it has a relatively independent evolutionary history. [3] Of the suggested 7 genera in the cormorant family, [3] the black-faced cormorant is part of the Old-Word cormorants Phalacrocorax. This genus diverged from its sister genus, the North-Pacific Cormorants Urile around 10 million years ago. [3]
The closest related species to the black-faced cormorant is the pied cormorant P. varius. [3]
Black-faced cormorants are endemic to the coastal regions of southern Australia and Tasmania. [8] The population is estimated to in the tens of thousands, [1] and is distributed across Western Australia, South Australia, Victoria and Tasmania, particularly in the Bass Strait and Spencer Gulf. [9] There are two independent populations: one on the southern coast of Western Australia, and the other on the coasts of Southern Australia, Victoria and Tasmania. [1]
Unlike the other cormorants found around the Australian continent, the habitat of the black-faced cormorant is exclusively coastal and marine. [7] They can be found in coastal waters, inlets, rocky shores, and offshore islands. [9] Occasionally black-faced cormorants can be found in estuaries of rivers. [4] Colonies are usually found on steep cliffs and rocky islands where they nest. [2] In the winter, black-faced cormorants leave coastal waters to breed in these colonies. [2]
Because the black-faced cormorant is an exclusively marine bird, its behaviours enable it to be well-adapted to its environment. It uses pursuit-diving to capture fish, [1] and nests high on cliffs [2] to avoid predators.
The black-faced cormorant eats fish almost exclusively. Its diet generally consists of benthic or demersal fish, as well as some cephalopods, and varies between breeding and non-breeding seasons. [2] It forages along the coast and at the mouths of rivers, [8] and will dive up to 12 m to catch its prey. [9] After diving, the black-faced cormorant will spread its wings to dry its feathers in the sun. [6] Black-faced cormorants will occasionally forage together in flocks. [6] While breeding, its preferred prey items are wrasse and silver trevally. [2] After the breeding season, silverbelly, wrasse, whiting and hardyheads are consumed most frequently. [2] Prey is consumed whole, and non-digestible parts are regurgitated in pellets. [2]
Breeding cormorants nest in pairs or breeding colonies on islands or cliffs. [4] They are likely monogamous like other cormorant species. [7] Nesting sites are on bare rocks, often close to water, and are solidly built of driftwood, seaweed, and other plants with a 35–45 cm diameter. [4] 2 to 3 elongated oval eggs are then laid. [4] The eggs are pale blue-green and measure 58 x 36 mm. [4]
The breeding season for black-faced cormorants varies between populations, and has been observed to occur anywhere from June to February. In the Bass Strait, the black-faced cormorant breeds from June to September, with a peak laying date around the end of July. [2] The reason for this may be to avoid hot summer temperatures that can affect chicks. [2] However, colonies in western and southern Australia breed later in the year, from September to February. [2]
Black-faced cormorants are considered 'Least Concern' by the IUCN, due to their large range and populations. [1] In Australia, they are considered 'Secure' federally as well as in South Australia, Western Australia, and Tasmania. [9] However, in Victoria, the black-faced cormorant is considered 'Vulnerable'. [9] Breeding colonies are very sensitive to human presences, [10] [2] so it is important that important breeding habitats are protected from disruption.
The double-crested cormorant is a member of the cormorant family of water birds. It is found near rivers and lakes and in coastal areas and is widely distributed across North America, from the Aleutian Islands in Alaska down to Florida and Mexico. Measuring 70–90 cm (28–35 in) in length, it is entirely black except for a bare patch of orange-yellow facial skin and some extra plumage that it exhibits in the breeding season when it grows a double crest in which black feathers are mingled with white. Five subspecies are recognized. It mainly eats fish and hunts by swimming and diving. Its feathers, like all cormorants, are not waterproof, and it must dry them out after spending time in the water. Once threatened by the use of DDT, the numbers of this bird have increased markedly in recent years.
The pelagic cormorant, also known as Baird's cormorant or violet-green cormorant, is a small member of the cormorant family Phalacrocoracidae. Analogous to other smallish cormorants, it is also called the pelagic shag occasionally. This seabird lives along the coasts of the northern Pacific; during winter it can also be found in the open ocean. Pelagic cormorants have relatively short wings due to their need for economical movement underwater, and consequently have the highest flight costs of any bird.
The little pied cormorant, little shag or kawaupaka is a common Australasian waterbird, found around the coasts, islands, estuaries, and inland waters of Australia, New Guinea, New Zealand, Timor-Leste, and Indonesia, and around the islands of the south-western Pacific and the subantarctic. It is a small short-billed cormorant usually black above and white below with a yellow bill and small crest, although a mostly black white-throated form predominates in New Zealand. Three subspecies are recognised. Until recently most authorities referred to this species as Phalacrocorax melanoleucos.
The great cormorant, known as the black shag or kawau in New Zealand, formerly also known as the great black cormorant across the Northern Hemisphere, the black cormorant in Australia, and the large cormorant in India, is a widespread member of the cormorant family of seabirds. The genus name is Latinised Ancient Greek, from φαλακρός and κόραξ, and carbo is Latin for "charcoal".
The European shag or common shag is a species of cormorant. It is the only member of the monotypic genus Gulosus. It breeds around the rocky coasts of western and southern Europe, southwest Asia and north Africa, mainly wintering in its breeding range except for the northernmost birds. In Britain this seabird is usually referred to as simply the shag. The scientific genus name derives from the Latin for glutton. The species name aristotelis commemorates the Greek philosopher Aristotle.
The Australian pied cormorant, also known as the pied cormorant, pied shag, or great pied cormorant, is a medium-sized member of the cormorant family. It is found around the coasts of Australasia. In New Zealand, it is usually known either as the pied shag or by its Māori name of kāruhiruhi. Older sources may refer to it as the "yellow-faced cormorant".
The spotted shag or pārekareka is a species of cormorant endemic to New Zealand. Though originally classified as Phalacrocorax punctatus, it is sufficiently different in appearance from typical members of that genus that for a time it was placed in a separate genus, Stictocarbo, along with a similar species, the Pitt shag. Subsequent genetic studies show that the spotted shag's lineage is nested within the typical shags.
The Indian cormorant or Indian shag is a member of the cormorant family. It is found mainly along the inland waters of the Indian Subcontinent but extends west to Sind and east to Thailand and Cambodia. It is a gregarious species that can be easily distinguished from the similar sized little cormorant by its blue eyes, small head with a sloping forehead and a long narrow bill ending in a hooked tip.
The little cormorant is a member of the cormorant family of seabirds. Slightly smaller than the Indian cormorant it lacks a peaked head and has a shorter beak. It is widely distributed across the Indian Subcontinent and extends east to Java, where it is sometimes called the Javanese cormorant. It forages singly or sometimes in loose groups in lowland freshwater bodies, including small ponds, large lakes, streams and sometimes coastal estuaries. Like other cormorants, it is often found perched on a waterside rock with its wings spread out after coming out of the water. The entire body is black in the breeding season but the plumage is brownish, and the throat has a small whitish patch in the non-breeding season. These birds breed gregariously in trees, often joining other waterbirds at heronries.
The Otago shag, together with the Foveaux shag formerly known as the Stewart Island shag and in its dark phase as the bronze shag, is a species of shag now found only in coastal Otago, New Zealand.
The guanay cormorant or guanay shag is a member of the cormorant family found on the Pacific coast of Peru and northern Chile. After breeding it spreads south to southern parts of Chile and north to Ecuador, and has also been recorded as far north as Panama and Colombia – probably a result of mass dispersal due to food shortage in El Niño years. Its major habitats include shallow seawater and rocky shores.
The little black cormorant is a member of the cormorant family of seabirds. It is common in smaller rivers and lakes throughout most areas of Australia and northern New Zealand, where it is known as the little black shag. It is around sixty centimetres long, and is all black with blue-green eyes.
The Cape cormorant or Cape shag is a bird endemic to the southwestern coasts of Africa.
The red-legged cormorant, also known as the red-legged shag, red-footed cormorant, red-footed shag, Gaimard's cormorant and grey cormorant, is a species of cormorant resident to the coastline of South America. It is the only member of the genus Poikilocarbo. It is non-colonial unlike most seabirds. The red-legged cormorant has not been observed wing-spreading, which is unusual among cormorant species.
The Chatham shag, also known as the Chatham Island shag, is a species of bird in the cormorant and shag family, Phalacrocoracidae. It is endemic to the Chatham Islands of New Zealand. For a long time the species was placed in the genus Phalacrocorax; today it is mostly placed with the other blue-eyed shags of New Zealand and Antarctica in the genus Leucocarbo. Its closest relative is the Otago shag of South Island.
The Kerguelen shag is a species of cormorant endemic to the Kerguelen Islands in the southern Indian Ocean, one of the most isolated places on Earth. Many authorities consider it a subspecies of the imperial shag.
The Heard Island shag, or Heard Island cormorant, is a marine cormorant native to the Australian territory comprising the Heard and McDonald Islands in the Southern Ocean, which is about 4,100 km south-west of Perth, Western Australia.
The Macquarie shag, Macquarie Island shag or Macquarie Island cormorant, is a marine cormorant native to Macquarie Island in the Southern Ocean, about halfway between Australia and Antarctica.
The Crozet shag, also known as the South Georgia cormorant, is a marine cormorant native to the Crozet, Prince Edward and Marion islands in the South Atlantic Ocean.
The Antarctic shag, sometimes referred to as the imperial cormorant, king cormorant, imperial shag, blue-eyed shag or Antarctic cormorant, is the only species of the cormorant family found in the Antarctic. It is sometimes considered conspecific with the Imperial shag.
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