Wild barley | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Clade: | Commelinids |
Order: | Poales |
Family: | Poaceae |
Subfamily: | Pooideae |
Genus: | Hordeum |
Species: | H. spontaneum |
Binomial name | |
Hordeum spontaneum (K. Koch) Thell. [1] | |
Synonyms | |
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Hordeum spontaneum, commonly known as wild barley or spontaneous barley, is the wild form of the grass in the family Poaceae that gave rise to the cereal barley (Hordeum vulgare). Domestication is thought to have occurred on two occasions, first about ten thousand years ago in the Fertile Crescent and again later, several thousand kilometres further east.
Wild barley is an annual grass and is very similar in form to cultivated barley (Hordeum vulgare) but has slightly narrower leaves, longer stems, longer awns, a brittle rachis, a longer, more slender seed spike and smaller grains. [2] Characteristics of the wild plant that enhance its survival and dispersal include the brittle rachis (the central part of the seed head), which breaks when the grain is ripe, and the hulled seeds, which are arranged in two rows. In cultivated varieties, the rachis is more durable and the seeds are usually arranged in two or six rows. In the east, barley is usually grown for human consumption and the naked form of the grain is preferred, while in the west, the hulled form is mainly grown. It is used for animal feed and for the production of malt for brewing. [3]
Wild barley is native to North Africa, the Middle East, parts of the Indian subcontinent and south-west China. Its range includes Libya, Egypt, Israel, Lebanon, Jordan, Syria, Crete, Cyprus, Turkey, Iran, Iraq, Afghanistan, Armenia, Azerbaijan, Kyrgyzstan, Turkmenistan, Uzbekistan, Pakistan, India and the Sichuan and Xizang provinces of China. [4]
Wild barley is found in a wide range of locations varying from deserts to Mediterranean-type habitats and highland habitats, growing in both cultivated and uncultivated fields. It is exposed to many environmental stresses including high temperatures, drought and high soil salinity and there are local adaptations to the microclimates in which it grows. This wild plant is potentially a genetic resource useful for the breeding of stress-tolerant varieties of the cultivated crop. [5]
The domestication of barley probably occurred around 10,000 years ago in the Israel and Jordan region of the Fertile Crescent. [2] Barley grains have been found at a number of archaeological sites in this area and the wild barleys here are more molecularly similar to the cultivated gene pool than are any other wild populations. AFLP technology indicates single domestication of barley, not, as had previously been hypothesized, a series of domestications in different parts of its range such as in Ethiopia and in the Mediterranean region. [2] Further research using haplotype frequency in different geographic parts of the range led to the inference that there were at least two domestication events, one in the Fertile Crescent and another some 1,500 to 3,000 km (932 to 1,864 mi) further east. The former gave rise to the cultivars grown in Europe and America while the latter gave rise to the diversity of cultivated crops in central Asia and the Far East. [6]
In Palestine, the ground was prepared for the sowing of barley in December, the ground sown with barley in January, and the barley harvested in April. [7]
Wheat is a grass widely cultivated for its seed, a cereal grain that is a staple food around the world. The many species of wheat together make up the genus Triticum ; the most widely grown is common wheat. The archaeological record suggests that wheat was first cultivated in the regions of the Fertile Crescent around 9600 BC. Botanically, the wheat kernel is a caryopsis, a type of fruit.
Einkorn wheat can refer either to a wild species of wheat (Triticum) or to its domesticated form. The wild form is T. boeoticum, and the domesticated form is T. monococcum. Einkorn is a diploid species of hulled wheat, with tough glumes ('husks') that tightly enclose the grains. The cultivated form is similar to the wild, except that the ear stays intact when ripe and the seeds are larger. The domestic form is known as "petit épeautre" in French, "Einkorn" in German, "einkorn" or "littlespelt" in English, "piccolo farro" in Italian and "escanda menor" in Spanish. The name refers to the fact that each spikelet contains only one grain.
Hordeum is a genus of annual and perennial plants in the grass family. They are native throughout the temperate regions of Africa, Eurasia, and the Americas.
Emmer wheat or hulled wheat is a type of awned wheat. Emmer is a tetraploid. The domesticated types are Triticum turgidum subsp. dicoccum and T. t. conv. durum. The wild plant is called T. t. subsp. dicoccoides. The principal difference between the wild and the domestic forms is that the ripened seed head of the wild plant shatters and scatters the seed onto the ground, while in the domesticated emmer, the seed head remains intact, thus making it easier for people to harvest the grain.
Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants, takes over control and care of another species, such as sheep or fungi, to obtain from them a steady supply of resources, such as meat, milk, or labor. The process is gradual and geographically diffuse, based on trial and error.
The Neolithic Revolution, also known as the First Agricultural Revolution, was the wide-scale transition of many human cultures during the Neolithic period in Afro-Eurasia from a lifestyle of hunting and gathering to one of agriculture and settlement, making an increasingly large population possible. These settled communities permitted humans to observe and experiment with plants, learning how they grew and developed. This new knowledge led to the domestication of plants into crops.
The founder crops or primary domesticates are a group of flowering plants that were domesticated by early farming communities in Southwest Asia and went on to form the basis of agricultural economies across Eurasia. As originally defined by Daniel Zohary and Maria Hopf, they consisted of three cereals, four pulses, and flax. Subsequent research has indicated that many other species could be considered founder crops. These species were amongst the first domesticated plants in the world.
Common wheat, also known as bread wheat, is a cultivated wheat species. About 95% of wheat produced worldwide is common wheat; it is the most widely grown of all crops and the cereal with the highest monetary yield.
During 10,000 years of cultivation, numerous forms of wheat, many of them hybrids, have developed under a combination of artificial and natural selection. This diversity has led to much confusion in the naming of wheats. Genetic and morphological characteristics of wheat influence its classification; many common and botanical names of wheat are in current use.
The Eastern Agricultural Complex in the woodlands of eastern North America was one of about 10 independent centers of plant domestication in the pre-historic world. Incipient agriculture dates back to about 5300 BCE. By about 1800 BCE the Native Americans of the woodlands were cultivating several species of food plants, thus beginning a transition from a hunter-gatherer economy to agriculture. After 200 BCE when maize from Mexico was introduced to the Eastern Woodlands, the Native Americans of the eastern United States and adjacent Canada slowly changed from growing local indigenous plants to a maize-based agricultural economy. The cultivation of local indigenous plants other than squash and sunflower declined and was eventually abandoned. The formerly domesticated plants returned to their wild forms.
Triticeae is a botanical tribe within the subfamily Pooideae of grasses that includes genera with many domesticated species. Major crop genera found in this tribe include wheat, barley, and rye; crops in other genera include some for human consumption, and others used for animal feed or rangeland protection. Among the world's cultivated species, this tribe has some of the most complex genetic histories. An example is bread wheat, which contains the genomes of three species with only one being a wheat Triticum species. Seed storage proteins in the Triticeae are implicated in various food allergies and intolerances.
Hordeum jubatum, with common names foxtail barley, bobtail barley, squirreltail barley, and intermediate barley, is a perennial plant species in the grass family Poaceae. It occurs wild mainly in northern North America and adjacent northeastern Siberia. However, as it escaped often from gardens it can be found worldwide in areas with temperate to warm climates, and is considered a weed in many countries. The species is a polyploid and originated via hybridization of an East Asian Hordeum species with a close but extinct relative of Californian H. brachyantherum. It is grown as an ornamental plant for its attractive inflorescences and when done flowering for its inflorescence.
Hordeum murinum is a species of flowering plant in the grass family Poaceae, commonly known as wall barley or false barley. It is a close relative of cultivated barley.
Hordeum pusillum, also known as little barley, is an annual grass native to most of the United States and southwestern Canada. It arrived via multiple long-distance dispersals of a southern South American species of Hordeum about one million years ago. Its closest relatives are therefore not the other North American taxa like meadow barley or foxtail barley, but rather Hordeum species of the Pampas of central Argentina and Uruguay. It is less closely related to the Old World domesticated barley, from which it diverged about 12 million years ago. It is diploid.
A crop wild relative (CWR) is a wild plant closely related to a domesticated plant. It may be a wild ancestor of the domesticated (cultivated) plant or another closely related taxon.
Barley, a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikelets and making it much easier to harvest. Its use then spread throughout Eurasia by 2000 BC. Barley prefers relatively low temperatures and well-drained soil to grow. It is relatively tolerant of drought and soil salinity but is less winter-hardy than wheat or rye.
Bere, pronounced "bear," is a six-row barley cultivated mainly on 5-15 hectares of land in Orkney, Scotland. It is also grown in Shetland, Caithness and on a very small scale by a few crofters on some of the Western Isles, such as North Uist, Benbecula, South Uist, Islay and Barra. It is probably Britain's oldest cereal in continuous commercial cultivation.
South and West Asia consists of a wide region extending from the present-day country of Turkey in the west to Bangladesh and India in the east.
In agriculture, shattering is the dispersal of a crop's seeds upon their becoming ripe. From an agricultural perspective this is generally an undesirable process, and in the history of crop domestication several important advances have involved a mutation in a crop plant that reduced shattering—instead of the seeds being dispersed as soon as they were ripe, the mutant plants retained the seeds for longer, which made harvesting much more effective. Non-shattering phenotype is one of the prerequisites for plant breeding especially when introgressing valuable traits from wild varieties of domesticated crops.
Ancient grains is a marketing term used to describe a category of grains and pseudocereals that are purported to have been minimally changed by selective breeding over recent millennia, as opposed to more widespread cereals such as corn, rice and modern varieties of wheat, which are the product of thousands of years of selective breeding. Ancient grains are often marketed as being more nutritious than modern grains, though their health benefits over modern varieties have been disputed by some nutritionists.