Long-tailed nightjar

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Long-tailed nightjar
Long-tailed nightjar (Caprimulgus climacurus climacurus) male.jpg
Male C. c. climacurus
Gambia
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Clade: Strisores
Order: Caprimulgiformes
Family: Caprimulgidae
Genus: Caprimulgus
Species:
C. climacurus
Binomial name
Caprimulgus climacurus
Vieillot, 1824

The Long-tailed Nightjar (Caprimulgus climacurus) is one of 98 species of nightjar in the family Caprimulgidae, the "true nightjars". It is a nocturnal, insectivorous bird characterized by its distinctive long tail. It is found throughout mainland Africa in open areas, arid semi-deserts, and savannas, with common sightings alongside roadways.

Contents

Description

The Long-tailed Nightjar ranges from about 29-43 cm in size and is sexually dimorphic. [2] It has a long, graduated tail, with its size ranging from 200-304 mm in males and 156-256 mm in females. [2] Its plumage is primarily rufous-brown to gray-brown in colour, but has pale bands (white in males, buff in females) across the forewing, as well as along the trailing edge of the flight feathers. [3] [4] The males can also have a white throat patch and brown bills, legs and feet. [5] A sexually immature Long-tailed Nightjar looks more similar to a female, however its plumage tends to be paler. [5]

The identification characteristics of the Long-tailed Nightjar are similar to those of the Slender-tailed Nightjar (Caprimulgus clarus) and the Square-tailed Nightjar (Caprimulgus fossii). Specifically, when the Long-tailed Nightjar is non-breeding or moulting, their tails can be shorter. [4] However, these species can be distinguished as they tend to occur in non-overlapping geographic areas and also have different songs. [2] The song of the Long-tailed Nightjar is composed of a rapid, higher-pitched and sustained churring, at a frequency of about 42-43 units per second, which is much quicker than both the Slender-tailed Nightjar (7-8.6 units per second) and the Square-tailed Nightjar (16-18 units per second). [2] In regions where the Slender-tailed Nightjar's range geographically overlaps with that of the Long-tailed Nightjar, the two can be distinguished as the Long-tailed Nightjar's plumage is relatively more red in colour. [4]

The moulting pattern in which the Long-tailed Nightjar replaces its primary feathers is a typical strategy amongst nightjars, where the primaries are replaced in a descendant manner. [6] However, its secondaries moult in a relatively unusual manner, with two moult centres. [6] One moult centre begins with the outermost secondary feather and moults in an ascendant manner towards the fifth secondary, whereas the second moult centre begins with the ninth secondary and follows a centrifugal moult pattern (outermost to innermost moulting). [6] The moult duration has been estimated to take approximately 70 days. [7] The moult tends to arrest in the mid-winter. [2]

Taxonomy and Systematics

Diversity within nightjar species, including the Long-tailed Nightjar, tends to be high both genetically and morphologically, resulting in many subspecies. [8] There are three recognized geographically distinct subspecies of Long-tailed Nightjar, C. c. climacurus, C. c. sclateri and C. c. nigricans. [2] The subspecies C. c. climacurus can be found from Mauritania to Sudan, eastern Congo and western Ethiopia. [9] C. c. sclateri can be found from Guinea to northwestern Uganda. [9] C. c. nigricans can be found in eastern Sudan and Ethiopia in the Nile Valley. [9] C. c. sclateri has relatively darker and redder plumage, with its tail being dark brown on top and black below. [2] C. c. nigricans has overall very dark plumage, with distinct black colouration on its upperparts interpolated with white specks. [2] Recent reconstructions of the nightjars' phylogeny indicate that the Long-tailed Nightjar is most closely related to the Square-tailed Nightjar. [10] C. climacurus, alongside C. fossii and C. clarus, were previously classified under a separate genus, known as Scotornis. [11]

Habitat and Distribution

The long-tailed nightjar is found in the Afrotropical realm [12] in multiple African countries including Angola, Benin, Burkina Faso, Cameroon, Central African Republic, Chad, Republic of the Congo, Democratic Republic of the Congo, Ivory Coast, Eritrea, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Kenya, Liberia, Mali, Mauritania, Niger, Nigeria, Senegal, Sierra Leone, Sudan, Tanzania, Togo and Uganda. [2] [4] It is found mainly in open areas of the northern Afrotropics. [2] This includes roadsides, savannas, semi-arid deserts and lowland forest clearings. [5] It can often be found in the company of the Standard-winged Nightjar (Macrodipteryx longipennis). [13]

Behaviour and Ecology

Reproduction

The laying season of the Long-tailed Nightjar varies depending on geographic location. [2] The laying season is from March to September in Gambia and Senegal, May to August in northern Nigeria, January to October in southern Nigeria, March to August in Mali, March to June in northeastern Congo and February to April or April to June in eastern Africa. [2]

Long-tailed Nightjar amongst leaf-litter with its eyes closed to slits. Long-tailed Nightjar - Gambia (32651080875).jpg
Long-tailed Nightjar amongst leaf-litter with its eyes closed to slits.

No nest is built for the eggs, rather a clear patch of land, approximately 4 x 5 cm in size, is chosen. [2] Nest locations include bare land, open soil, sand, clear patches between pebbles or leaf-litter, beside pathways, in a thicket or under small bushes in fields. [2] The clutch size is typically two eggs. [2] The eggs are ellipsoidal in shape, matte in finish and colouration varying from whitish, cream, buff or pinkish - typically matching the ground colour. [2] Additionally, grey, grey-purple and chestnut brown blotches and thick marbling fill in the egg's surface, further aiding with camouflage [2] .The incubating bird tends to rest flat on top of the eggs with its eyelids closed. [2]

Diet and Feeding

Similar to other nightjars, the diet of the Long-tailed Nightjar predominantly consists of insects. These insects include grasshoppers, ants, beetles, moths and winged termites. [5] The most common feeding grounds for the Long-tailed Nightjar are areas near cattle trails, horse trails, camel trails, farmland and generally open spaces. [12] Often, it perches on the ground and quickly flies up to catch insect that pass through its visual field. [12] It has also been found to forage over open water or marshland, typically drinking water beforehand. [12] It drinks water by hovering at progressively lower altitudes until it is about 20 cm above the water surface, where it soon after drops to the surface, dips its beak and flies away. [12] When hunting over water or amongst trees, it sustains flight in a straight line, then abruptly swerves or dives for its prey. [12] The Long-tailed nightjar will usually consume its prey while perched on the ground, and will at times exceptionally consume terrestrial (rather than aerial) insects. The Long-tailed Nightjar also shows an increase in twilight foraging activity near the new moon. [14]

Vocalization

Relative to other nightjars, the Long-tailed Nightjar's call is a faster, higher-pitched, prolonged trill. [4] During flight, especially in the process of catching prey, the Long-tailed nightjar often produces a "chuck" or "chiow". [4] [12]

Status and Conservation

When illuminated by flashlights or automobile headlights, the eyes of Long-tailed Nightjars distinctively appear red. Caprimulgus fossii subsp mossambicus, Pretoriuskop, Birding Weto, a.jpg
When illuminated by flashlights or automobile headlights, the eyes of Long-tailed Nightjars distinctively appear red.

The Long-tailed Nightjar is listed as least concern according to the IUCN Red List. [15] Despite the population size being unknown, it occupies a large range, with its extent of occurrence being estimated at 9 770 000 km2. [15] However, human activity, especially road traffic, has been a major mortality factor for the Long-tailed Nightjar. [16] The Long-tailed Nightjar has been found to frequent roads at dusk and night. [16] With increasing road-traffic comes increasing road-related mortality, as the Long-tailed nightjar has often been observed as unable to fly out of the way of approaching vehicles in time to escape being struck. [16] Recommended actions to lower their road mortality includes periodically honking the horn, slowing down speeds and flashing the headlights of vehicles. [16]

Related Research Articles

<span class="mw-page-title-main">Nightjar</span> Family of birds

Nightjars are medium-sized nocturnal or crepuscular birds in the family Caprimulgidae and order Caprimulgiformes, characterised by long wings, short legs, and very short bills. They are sometimes called bugeaters, their primary source of food being insects. Some New World species are called nighthawks. The English word nightjar originally referred to the European nightjar.

<span class="mw-page-title-main">European nightjar</span> Migratory bird found in Eurasia and Africa

The European nightjar, common goatsucker, Eurasian nightjar or just nightjar, is a crepuscular and nocturnal bird in the nightjar family that breeds across most of Europe and the Palearctic to Mongolia and Northwestern China. The Latin generic name refers to the old myth that the nocturnal nightjar suckled from goats, causing them to cease to give milk. The six subspecies differ clinally, the birds becoming smaller and paler towards the east of the range. All populations are migratory, wintering in sub-Saharan Africa. Their densely patterned grey and brown plumage makes individuals difficult to see in the daytime when they rest on the ground or perch motionless along a branch, although the male shows white patches in the wings and tail as he flies at night.

<span class="mw-page-title-main">Nighthawk</span> Subfamily of birds

The nighthawk is a nocturnal bird of the subfamily Chordeilinae, within the nightjar family, Caprimulgidae, native to the western hemisphere. The term "nighthawk", first recorded in the King James Bible of 1611, was originally a local name in England for the European nightjar. Its use in the Americas to refer to members of the genus Chordeiles and related genera was first recorded in 1778.

<span class="mw-page-title-main">Standard-winged nightjar</span> Species of bird

The standard-winged nightjar is a nocturnal bird in the nightjar family.

<span class="mw-page-title-main">Jerdon's nightjar</span> Species of bird

Jerdon's nightjar is a medium-sized nightjar species native to southern India and Sri Lanka. Formerly considered as a subspecies of the long-tailed nightjar, it is best recognized by its distinctive call which sounds like a wooden plank being beaten periodically with each note ending in a quaver. The common name commemorates Thomas C. Jerdon who described the species.

<i>Caprimulgus</i> Genus of birds

Caprimulgus is a large and very widespread genus of nightjars, medium-sized nocturnal birds with long pointed wings, short legs and short bills. Caprimulgus is derived from the Latin capra, "nanny goat", and mulgere, "to milk", referring to an old myth that nightjars suck milk from goats. The common name "nightjar", first recorded in 1630, refers to the nocturnal habits of the bird, the second part of the name deriving from the distinctive churring song.

<span class="mw-page-title-main">Sykes's nightjar</span>

The Sykes's nightjar, Sykes' nightjar, Sind nightjar or Sindh nightjar is a nightjar species found in northwestern South Asia. They are insectivorous nocturnal birds known for their sandy plumage used for camouflage and their excellent flight abilities, allowing them to catch their prey mid-air. This species has a large range with stable population size and therefore has been listed as “Least Concern” with no prediction of substantial human influence on their habitats in the near future.

<span class="mw-page-title-main">Savanna nightjar</span> Species of bird

The savanna nightjar, also known as allied nightjar or Franklin's nightjar, is a species of nightjar found in South and Southeast Asia. The IUCN Red List has assessed the species to be of least concern because it has a large range and its population trend is stable. As other nightjars, this nocturnal bird is characterised by its large eyes, gaping mouth and excellent camouflage. It can be distinguished from similar southasian nightjar species, like the chirruping nightjar, by its unique vocalisations.

<span class="mw-page-title-main">Eared nightjar</span> Subfamily of birds

The eared nightjars are a small group of nocturnal birds in the nightjar family, although the taxonomy is uncertain. There are seven species, mainly found in forest and scrub from China to Australia. Five species are placed in the genus Eurostopodus, the other two species in Lyncornis. They are long winged birds with plumage patterned with grey and brown to camouflage them when resting on the ground. They feed on insects caught in flight. A single white egg is laid directly on the ground and incubated by both adults. The chicks can walk soon after hatching.

<span class="mw-page-title-main">Bates's nightjar</span> Species of bird

Bates's nightjar or the forest nightjar, is a bird species of the family Caprimulgidae, found in the rainforests of western Sub-Saharan Africa.

<span class="mw-page-title-main">Square-tailed nightjar</span> Species of bird

The square-tailed nightjar is a species of nightjar in the family Caprimulgidae which is native to tropical and subtropical woodlands of the Afrotropics. It has an extensive range south of the African equator. Despite not having a completely "square tail", its naming highlights a distinguishing field mark. The similarly plumaged slender-tailed nightjar, found in dry bush country of the tropics, differs namely by its protruding central tail feathers. It is alternatively known as the Gabon nightjar or Gaboon nightjar or the Mozambique nightjar.

<span class="mw-page-title-main">Band-winged nightjar</span> Species of bird

The band-winged nightjar or greater band-winged nightjar is a species of nightjar in the family Caprimulgidae. It is widespread in South America, where it is found in the Andes, Venezuelan Coastal Range, Santa Marta Mountains, Tepuis, most of Chile, Argentina, Paraguay, Uruguay, and eastern Brazil. It occurs in a wide range of habitats, from the edge of humid montane forests to shrubby semi-deserts and urban rooftops.

<span class="mw-page-title-main">Large-tailed nightjar</span> Species of bird

The large-tailed nightjar is a species of nightjar known for its distinctive long, broad tail and cryptic plumage that allows it to blend into its surroundings. Found along the southern Himalayan foothills, eastern South Asia, Southeast Asia and northern Australia, its natural habitats are subtropical or tropical moist lowland forest, subtropical or tropical mangrove forest, and subtropical or tropical moist montane forest.

<span class="mw-page-title-main">Fiery-necked nightjar</span> Species of bird

The fiery-necked nightjar is a species of nightjar in the family Caprimulgidae, which is found mostly in Africa south of the equator, though it has been spotted in a few countries north of the equator. It is most often found in woodland savannas or other deciduous woodlands. It is usually distinguished by its tawny coloured collar which gives the species its common name. It has a distinctive call that many have rendered as 'good-lord-deliver-us'. The fiery-necked nightjar is an insectivorous species that mostly eats butterflies, moths and other insects. The fiery-necked nightjar breeds after the dry season and typically produce two clutches with two eggs per clutch.

<span class="mw-page-title-main">Montane nightjar</span> Species of bird

The montane nightjar, mountain nightjar or Abyssinian nightjar, is a species of nightjar in the family Caprimulgidae. It is native to upland regions of Central and Eastern Africa where it is a locally common species.

<span class="mw-page-title-main">Rufous-cheeked nightjar</span> Species of bird

The rufous-cheeked nightjar is a species of nightjar in the family Caprimulgidae. It is an intra-African migrant that breeds in the south of its range. It spends the non-breeding season in eastern Nigeria, Cameroon, southern Chad and Sudan, the Republic of the Congo, the D.R.C. and western Central African Republic. Throughout the year it can be found in Southern Africa. It is a nocturnal species that feeds on flying insects, mainly hunting at dusk and the early night. They can hunt at night thanks to their large eyes that are adept at seeing in the dark. They cannot see in complete darkness and most likely cannot see colours either. The rufous-cheeked nightjar forms monogamous pairs and the egg laying period is from September to November. It looks very similar to other African nightjars and they are frequently misidentified during research and studies.

<span class="mw-page-title-main">Freckled nightjar</span> Species of bird

The freckled nightjar or freckled rock nightjar is a species of nightjar in the family Caprimulgidae. It has a wide yet patchy distribution throughout the Afrotropics.

<span class="mw-page-title-main">White-winged nightjar</span> Species of bird

The white-winged nightjar is a species of nightjar in the family Caprimulgidae. It is found in Bolivia, Brazil and Paraguay.

<span class="mw-page-title-main">Spotted nightjar</span> Species of bird

The spotted nightjar or spotted eared-nightjar is a species of nightjar in the family Caprimulgidae. It inhabits much of mainland Australia and has also been found in several Indonesian islands. Its natural habitats are open forests and woodlands, scrub, spinifex and tussock grassland, savannah woodland and mangroves.

<span class="mw-page-title-main">Grey nightjar</span> Species of bird

The grey nightjar is a species of nightjar found in East Asia, breeding from southeastern Russia south through China, the Korean Peninsula and Japan to northern Indochina and westward along the Himalayas. It is largely migratory, wintering in Indochina south to Java in Indonesia, but is resident in warmer areas in the south of its breeding range. It is sometimes treated as a subspecies of the jungle nightjar, its South Asian relative. The grey nightjar breeds and forages in early successional habitats surrounded by forests; its populations have declined since the 1970s in Japan. Like all nightjars, it is crepuscular to nocturnal, feeding on flying insects, including moths, beetles, flying ants, grasshoppers, and others.

References

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  8. Sigurðsson, Snorri; Cracraft, Joel (March 2014). "Deciphering the diversity and history of New World nightjars (Aves: Caprimulgidae) using molecular phylogenetics: Phylogeny of New World Nightjars". Zoological Journal of the Linnean Society. 170 (3): 506–545. doi:10.1111/zoj.12109.
  9. 1 2 3 The Clements Checklist of Birds of the World.[ page needed ]
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  11. Han, Kin-Lan (2006). Molecular systematics of nightjars and nighthawks (Caprimulgidae) (Thesis). ProQuest   305298963.[ page needed ]
  12. 1 2 3 4 5 6 7 Jackson, Hd (September 2003). "A review of foraging and feeding behaviour, and associated anatomical adaptations, in Afrotropical nightjars". Ostrich. 74 (3–4): 187–204. Bibcode:2003Ostri..74..187J. doi:10.2989/00306520309485392.
  13. Borrow, Nik; Demey, Ron (2023). Field Guide to Birds of Senegal and The Gambia: Second Edition. Bloomsbury USA. ISBN   978-1-3994-0220-0.[ page needed ]
  14. Ashdown, Robert A. M.; McKechnie, Andrew E. (October 2008). "Environmental correlates of Freckled Nightjar (Caprimulgus tristigma) activity in a seasonal, subtropical habitat". Journal of Ornithology. 149 (4): 615–619. Bibcode:2008JOrni.149..615A. doi:10.1007/s10336-008-0309-7.
  15. 1 2 "Long-tailed Nightjar (Caprimulgus climacurus) - BirdLife species factsheet". datazone.birdlife.org. Retrieved 2024-10-14.
  16. 1 2 3 4 Jackson, H.D.; Slotow, R. (September 2002). "A review of Afrotropical nightjar mortality, mainly road kills". Ostrich. 73 (3–4): 147–161. Bibcode:2002Ostri..73..147J. doi:10.1080/00306525.2002.11446745.