Pachyaena Temporal range: | |
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Pachyaena gigantea jaws | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | † Mesonychia |
Family: | † Mesonychidae |
Genus: | † Pachyaena Cope, 1874 |
Type species | |
†Pachyaena ossifraga Cope, 1874 | |
Other species | |
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Pachyaena (literally, "thick hyena") was a genus of heavily built, relatively short-legged mesonychids, early Cenozoic mammals that evolved before the origin of either modern hoofed animals or carnivores, and combined characteristics similar to both. The genus likely originated from Asia and spread to Europe, and from there to North America across a land bridge in what is now the North Atlantic ocean. [1] Various described species of Pachyaena ranged in size from a coyote to a bear. However, a 2007 paper suggests that Pachyaena may be paraphyletic, with P. ossifraga being closer to Synoplotherium , Harpagolestes and Mesonyx than to P. gigantea. [2]
Unlike many mesonychids, Pachyaena is known from skeletal material in addition to skulls and jaws. Analysis of three wolf- to bear-sized species from the early Eocene of Wyoming (Willwood formation) indicates they all had many adaptations for running, including paraxonic compressed feet with a vestigial first digit, lower sections of the limbs elongated compared with the upper sections, and limb joints with movement mostly restricted to a sagittal plane (back-and-forth movement). All these characteristics are common to both ungulates that run to escape predators and carnivores that run to pursue prey, though they probably evolved independently in mesonychids. Some adaptations are more typical of the grade of cursorial carnivores; others are more specialized, as in ungulates. Pachyaena was likely built for endurance rather than speed; the overall body shape of the genus would have resembled a modern tapir. [3]
Size varied depending on the species: For P. gracilis, the body mass estimated based in femur length is 24.0 kg, for P. ossifraga is about 56.9 kg, and for the large P. gigantea is 81.7 kg. [4] [5] However, weight estimations of P. gigantea vary from 129 to 396 kg. [6] Using Zhou's equation (Unpublished thesis, 1995) of ln(body mass) = 1.327 * ln(m2 length * width) -3.355, a body mass of 130 kg is calculated for P. gigantea.
The genus' greatest diversity is seen in North America, with P. ossifraga known from Latest Paleocene strata of Wyoming, and P. gigantea, P. intermedia, and P. gracilis known from Early Eocene strata of Wyoming. P. nemegetica is known from Late Paleocene strata of Mongolia.
Andrewsarchus, meaning "Andrews' ruler", is an extinct genus of artiodactyl that lived during the Middle Eocene in what is now China. The genus was first described by Henry Fairfield Osborn in 1924 with the type species A. mongoliensis based on a largely complete cranium. A second species, A. crassum, was described in 1977 based on teeth. A mandible, formerly described as Paratriisodon, does probably belong to Andrewsarchus as well. The genus has been historically placed in the families Mesonychidae or Arctocyonidae, or was considered to be a close relative of whales. It is now regarded as the sole member of its own family, Andrewsarchidae, and may have been related to entelodonts. Fossils of Andrewsarchus have been recovered from the Middle Eocene Irdin Manha, Lushi, and Dongjun Formations of Inner Mongolia, each dated to the Irdinmanhan Asian land mammal age.
Creodonta is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ancestral to the modern Carnivora, this order is now usually considered a polyphyletic assemblage of two different groups, the oxyaenids and the hyaenodonts, not a natural group. Oxyaenids are first known from the Palaeocene of North America, while hyaenodonts hail from the Palaeocene of Africa.
Mesonychia is an extinct taxon of small- to large-sized carnivorous ungulates related to artiodactyls. Mesonychians first appeared in the early Paleocene, went into a sharp decline at the end of the Eocene, and died out entirely when the last genus, Mongolestes, became extinct in the early Oligocene. In Asia, the record of their history suggests they grew gradually larger and more predatory over time, then shifted to scavenging and bone-crushing lifestyles before the group became extinct.
Phenacodus is an extinct genus of mammals from the late Paleocene through middle Eocene, about 55 million years ago. It is one of the earliest and most primitive of the ungulates, typifying the family Phenacodontidae and the order Perissodactyla.
Miacis is an extinct genus of placental mammals from clade Carnivoraformes, that lived in North America from the early to middle Eocene.
Mesonyx is a genus of extinct mesonychid mesonychian mammal. Fossils of the various species are found in Early to Late Eocene-age strata in the United States and Early Eocene-aged strata in China, 51.8—51.7 Ma (AEO).
Chriacus is an extinct genus of placental mammals that lived in what is now North America during the Paleocene epoch and died out after the early Eocene. In life, members of the genus would have looked something like a kinkajou or binturong, though they were not closely related to any living mammal. Well preserved fossils allow clear information on what they looked like. They were about 1 metre (3.3 ft) long including a long, robust tail, which may or may not have been prehensile. Other features include a light build, weighing approximately 7 kg (15 lb), and many adaptations typical of animals that live in trees. These include walking on the soles of their five-toed feet, and having long, curved, compressed claws. The powerfully built limbs had flexible joints, especially the ankles, an adaptation that allows an animal to turn its hind feet behind it, like modern tree squirrels, in order to climb downward. They were probably omnivores, eating fruit, eggs, insects and small mammals.
Arctocyon is an extinct genus of large placental mammals, part of the possibly polyphyletic family Arctocyonidae. The type species is A. primaevus, though up to five other species may be known. Fossils of Arctocyon have been found in Europe and North America. Arctocyon was originally named as a subgenus of the bear-dog Amphicyon, though was subsequently found to belong to a genus and family of its own. The relationship between arctocyonids and other placentals is unclear, with early classification efforts placing them as carnivores or creodonts, and later ones classifying them under Condylarthra, a wastebasket taxon for various early ungulates. More recent phylogenetic analyses suggest that arctocyonids are an artificial assemblage of several distantly-related placental lineages, and Arctocyon may be closely related to Loxolophus and to pantodonts.
Oxyaena is an extinct genus of placental mammals from extinct subfamily Oxyaeninae within extinct family Oxyaenidae, that lived in Europe, Asia and North America during the early Eocene.
Mesonychidae is an extinct family of small to large-sized omnivorous-carnivorous mammals. They were endemic to North America and Eurasia during the Early Paleocene to the Early Oligocene, and were the earliest group of large carnivorous mammals in Asia. Once considered a sister-taxon to artiodactyls, recent evidence now suggests no close connection to any living mammal. Mesonychid taxonomy has long been disputed and they have captured popular imagination as "wolves on hooves", animals that combine features of both ungulates and carnivores. Skulls and teeth have similar features to early whales, and the family was long thought to be the ancestors of cetaceans. Recent fossil discoveries have overturned this idea; the consensus is that whales are highly derived artiodactyls. Some researchers now consider the family a sister group either to whales or to artiodactyls, close relatives rather than direct ancestors. Other studies define Mesonychia as basal to all ungulates, occupying a position between Perissodactyla and Ferae. In this case, the resemblances to early whales would be due to convergent evolution among ungulate-like herbivores that developed adaptations related to hunting or eating meat.
Dissacus is a genus of extinct carnivorous jackal to coyote-sized mammals within the family Mesonychidae, an early group of hoofed mammals that evolved into hunters and omnivores. Their fossils in Paleocene to Early Eocene aged strata in France, Asia and southwest North America, from 66 to 50.3 mya, existed for approximately 15.7 million years.
Hapalodectidae is an extinct family of relatively small-bodied mesonychian placental mammals from the Paleocene and Eocene of North America and Asia. Hapalodectids differ from the larger and better-known mesonychids by having teeth specialized for cutting, while the teeth of other mesonychids, such as Mesonyx or Sinonyx, are more specialized for crushing bones. Hapalodectids were once considered a subfamily of the Mesonychidae, but the discovery of a skull of Hapalodectes hetangensis showed additional differences justifying placement in a distinct family. In particular, H. hetangensis has a postorbital bar closing the back of the orbit, a feature lacking in mesonychids. The skeleton of hapalodectids is poorly known, and of the postcranial elements, only the humerus has been described. The morphology of this bone indicates less specialization for terrestrial locomotion than in mesonychids.
Ankalagon saurognathus is an extinct carnivorous mammal of the family Mesonychidae, endemic to North America during the Paleocene epoch, existing for approximately 3.1 million years.
Didymictis is an extinct genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America and Europe from the late Paleocene to middle Eocene.
Psittacotherium is an extinct genus of taeniodonts from extinct tribe Psittacotheriini within subfamily Stylinodontinae and family Stylinodontidae, that lived in North America from early to late Paleocene. With a weight between 35.6 kg (78 lb) and 71.2 kg (157 lb), and length of 1.125 m, it had similar size of a large dog.
Tillodontia is an extinct suborder of eutherian mammals known from the Early Paleocene to Late Eocene of China, the Late Paleocene to Middle Eocene of North America where they display their maximum species diversity, the Middle Eocene of Pakistan, and the Early Eocene of Europe. Leaving no descendants, they are most closely related to the pantodonts, another extinct group. The tillodonts were medium- to large-sized animals that probably fed on roots and tubers in temperate to subtropical habitats.
Harpagolestes is an extinct genus of hyena like, bear sized mesonychid mesonychian that lived in Central and Eastern Asia and western and central North America during the middle to late Eocene. It has been suggested that Harpagolestes is a wastebasket genus for fragmentary Eocene mesonychids.
Palaeonictis is an extinct genus of placental mammals from extinct subfamily Palaeonictinae within extinct family Oxyaenidae, that lived in Europe and North America from the late Paleocene to the early Eocene.
Apatemys is a member of the family Apatemyidae, an extinct group of small and insectivorous placental mammals that lived in the Paleogene of North America, India, and Europe. While the number of genera and species is less agreed upon, it has been determined that two apatemyid genera, Apatemys and Sinclairella, existed sequentially during the Eocene in North America. The genus Apatemys, living as far back as 50.3 million years ago (mya), existed through part of the Wasatchian and persisted through the Duchesnean, and Sinclairella followed, existing from the Duchesnean through the Arikareean. Examinations of specimens belonging to the genus Apatemys suggest adaptations characteristic of arboreal mammals.
Protictis is an extinct paraphyletic genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America from early Paleocene to middle Eocene.