Platycephalidae | |
---|---|
Sunagocia otaitensis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Scorpaeniformes |
Suborder: | Platycephaloidei |
Family: | Platycephalidae Swainson, 1839 [1] |
Genera [2] | |
The Platycephalidae are a family of marine ray-finned fish, most commonly referred to as flatheads. They are relatives of the popular lionfish, and belong to the order Perciformes.
Platycephalidae was first proposed as a family in 1839 by the English naturalist William John Swainson. [1] The 5th edition of Fishes of the World classifies this family within the suborder Platycephaloidei in the order Scorpaeniformes. [3] Other authorities differ and do not consider the Scorpaeniformes to be a valid order because the Perciformes is not monophyletic without the taxa within the Scorpaeniformes being included within it. These authorities consider the Platycephalidae to belong to the suborder Platycephaloidei, along with the families Bembridae, Parabembridae, Hoplichthyidae and Plectrogeniidae within the Perciformes. [4]
Platycephalidae has the following 17 genera (including about 86 species) classified within it: [2]
Platycephalidae has been divided into as many as 5 subfamilies by some authors but Fishes of the World does not recognise subfamilies but does state that some authors recognise two subfamilies. [3] The two subfamilies are Platycephalalinae, containing Elates and Platycephalus, and Onigociinae, containing the remaining genera. [5] These two groupings are thought to have become divergent in the Eocene with the Platycephalinae being predominantly temperate and Onigociinae, being predominantly tropical. The basal Platycephalinae species are confined to southern Australia and the more derived taxa have diversified in the tropical Indo-West Pacific. [6]
The genus Sorsogona is recognised by Fishbase but, apparently, it does not include the type species, Sorsogona serrulata, as a species within it. [7] Catalog of Fishes treats this genus as a synonym of Rogadius but classifies all the species in Ratabulus , including the type species, which it treats as a synonym of Ratabulus tuberculatus . [8] [9]
The name of the family is derived from the Greek words platy, meaning "flat", and kephale for "head". [10]
Platycephalids are small to medium-sized fish. Most species are small, reaching an average of 10 cm in length. However, a few species in the genus Platycephalus are known to grow up to over a 1 m (3 ft 3 in) in length. [11] Their most distinctive characteristic is the flattened shape of their heads. While the rest of their bodies is shaped similarly to other fish that inhabit the areas they frequent, their heads are triangle-shaped and dorsoventrally depressed, giving them the shape of a trowel or an artist's spatula. [12] Their eyes are situated on the upper surface, in accordance with their bottom-dwelling lifestyle. They possess two complete dorsal fins, the first one supported by six to 9 strong spines.
Platycephalidae can be found naturally in coastal waters throughout the Indo-Pacific. [3] A few species have been caught in the eastern Mediterranean, having traveled there from the Red Sea through the Suez Canal. [13]
Flatheads are mostly marine demersal fish, often resting directly on the seabed, sometimes partially buried in sand or mud. They can be found in a wide range of depths, ranging from 10 m to the edge of the continental shelf at depths of about 300 m (980 ft). [12]
Flatheads are carnivorous, feeding on small fish and crustaceans. They lie in wait buried by sand, with only their eyes poking out from the substrate. When prospective prey walks or swims close to the platycephalid's head, the flathead strikes rapidly, engulfing the prey in its large mouth. As flatheads are ambush predators they are expected to be relatively sedentary and not move large distances as adults. However, recent research has shown that part of the population of some flathead species makes long-distance movements or spawning migrations. [14] [15]
Active commercial fisheries are geared towards members of the family. In Japan, some species are the subject of experimental aquaculture programs. Flatheads are commonly caught on rod and line. [16] The larger species are considered game fish.
Platycephalus is a genus of mostly marine, demersal ray-finned fish belonging to the family Platycephalidae. They are found in the eastern Mediterranean, the Indian Ocean and western Pacific Ocean.
The tentacled flathead, also known as the Indian Ocean crocodilefish, Madagascar flathead or longhead flathead, is a species of marine ray-finned fish belonging to the family Platycephalidae, the flatheads. This species is in the western Indian Ocean, including the Red Sea and the Mediterranean, having invaded as a Lessepsian migrant through the Suez Canal. It is the only species in the monotypic genus Papilloculiceps.
Bembridae, the deep-water flatheads, are a family of bottom-dwelling ray-finned fishes. They are found in the Indian and western Pacific Oceans.
Platycephaloidei is a suborder of ray-finned fishes, part of the order Scorpaeniformes, and includes the flatheads, ghost flatheads and sea robins.
Hoplichthys, the ghost flatheads, is a genus of marine ray-finned fishes native to the Indian and Pacific Oceans. This genus is the only member of the family Hoplichthyidae.
Parabembras is a genus of marine ray-finned fish belonging to the family Bembridae, the deepwater flatheads, although they are sufficiently different from the other genera in that family to be classified as their own family, Parabembradidae, by some authorities. These fishes are found in the Indian Ocean and the western Pacific Ocean.
Cociella is a genus of marine ray-finned fishes belonging to the family Platycephalidae, the flatheads. These fishes are found in the Indo-Pacific region.
Cymbacephalus is a genus of marine ray-finned fishes belonging to the family Platycephalidae, the flatheads. These fishes are found in the Indo-Pacific region.
Grammoplites is a genus of marine ray-finned fishes belonging to the family Platycephalidae, the flatheads. These fishes are found in the Indo-Pacific region.
The spiny flathead, also known as the whitefin flathead or Bengal flathead, is a species of marine ray-finned fish belonging to the family Platycephalidae, the flatheads. It is found in the Indo-West Pacific region. This species is the only known member of its genus.
Onigocia is a genus of marine ray-finned fishes belonging to the family Platycephalidae, the flatheads. They are found in the Indo-Pacific region.
Ratabulus is a genus of marine, demersal ray-finned fish belonging to the family Platycephalidae. These fishes are native to the eastern Indian Ocean and the western Pacific Ocean.
Rogadius is a genus of marine, demersal ray-finned fish belonging to the family Platycephalidae. These fishes are native to the Indian Ocean and the western Pacific Ocean.
The Guinea flathead is a species of marine ray-finned fish belonging to the family Platycephalidae, the flatheads. It is found in the eastern Atlantic Ocean along the western coast of Africa. This species is the only known member of the genus Solitas. It is also the only flathead species which has a natural distribution which lies outside of the Indo-Pacific.
Insidiator is a genus of marine, demersal ray-finned fish belonging to the family Platycephalidae. These fishes are native to the eastern Indian Ocean and western Pacific Ocean.
Sunagociais a genus of marine, demersal ray-finned fish belonging to the family Platycephalidae. These fishes are native to the Indian and Pacific oceans.
Thysanophrys is a genus of marine, demersal ray-finned fish belonging to the family Platycephalidae. These fishes are native to the Indian Ocean and the western Pacific Ocean.
Plectrogenium, is a genus of marine ray-finned fishes, the stinger flatheads, the only genus classified within the subfamily Plectrogeninae, which in turn is classified within the family Scorpaenidae. This genus is found in the Indian and Pacific Oceans.
Cociella martingomoni is a species of marine ray-finned fish belonging to the family Platycephalidae, the flatheads. It is found in the Andaman Sea.
Inegocia japonica, the Japanese flathead, rusty flathead or smooth flathead, is a species of marine ray-finned fish belonging to the family Platycephalidae, the flatheads. It is found in the Indo-West Pacific region.
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