Quercus douglasii

Last updated

Blue oak
Large Blue Oak.jpg
A large blue oak in a pasture in Mariposa County, California
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Fagales
Family: Fagaceae
Genus: Quercus
Subgenus: Quercus subg. Quercus
Section: Quercus sect. Quercus
Species:
Q. douglasii
Binomial name
Quercus douglasii
Quercus douglasii range map.jpg
Natural range
Synonyms [3]
  • Quercus douglasii var. ransomii(Kellogg) Beissn.
  • Quercus gambeliiLiebm. [note 1]
  • Quercus oblongifolia var. brevilobataTorr.
  • Quercus ransomiiKellogg

Quercus douglasii, known as blue oak, is a species of oak endemic to California, common in the Coast Ranges and the foothills of the Sierra Nevada. [4] It is California's most drought-tolerant deciduous oak, [5] and is a dominant species in the blue oak woodland ecosystem. It is occasionally known as mountain oak and iron oak. [6] [7]

Contents

Description

Quercus douglasii is a medium-sized tree with sparse foliage, generally 6–20 meters (20–66 feet) tall, with a trunk 36–60 centimeters (1–2 ft) in diameter at breast height. [5] Trunks are typically solitary, but some trees have multiple trunks. [5] [8] The tallest recorded specimen was found in Alameda County, at 28.7 m (94 ft). [6] [9] The trees grow slowly, about 30 cm (12 in) per year. [10] Individual trees over 500 years old have been recorded. [11]

The bark is light gray with many medium-sized dark cracks. The blue-green leaves are tough and leathery, [4] deciduous, 4–10 cm (1+12–4 in) long, and entire or shallowly lobed. The acorns are 2–3 cm (341+14 in) long, [8] with a moderately sweet kernel, and mature in 6–7 months from pollination.

Q. douglasii is monoecious and wind-pollinated. Flower buds take a growing season to develop into catkins. [5] Blue oak pollen is severely allergenic. [12]

Taxonomy

The species is one of over 80 named after Scottish botanist David Douglas. The common name "blue oak" derives from the dark blue-green tint of its leaves. [2]

Taxonomically it is placed in the white oak group [13] (subgenus Quercus, section Quercus).

Ecology

Q. douglasii prefers dry to moist soil and plenty of sunlight. [10] Its sparse foliage allows more light to reach the ground, and young trees may grow for decades below their parents' canopies. [5] The species often co-habitates with gray pine (Pinus sabiniana), [5] [14] and is also found with interior live oak (Q. wislizeni), coast live oak (Q. agrifolia), valley oak (Q. lobata), Oregon white oak (Q. garryana), [5] and canyon live oak (Q. chrysolepis). [15] Natural hybrids between Q. douglasii and the related Q. lobata, Q. garryana, [5] and shrub live oak (Q. turbinella) often occur where the species grow together in the same area. Sources consider Quercus × alvordiana to be a hybrid of Q. douglasii and either Q. turbinella [16] or Q. john-tuckeri . [5]

Old-growth blue oak woodland may be one of the most widespread old-growth forest ecosystems remaining in the state after European colonization. [17]

Drought tolerance

Quercus douglasii is the most drought-tolerant of California's deciduous oaks. It has a smaller canopy than less drought-tolerant relatives, and invests proportionally more growth into roots rather than leaves throughout its life cycle. [5] The leathery blue-green leaves contribute to its drought resistance; [4] during drought, the leaf color is more pronounced. Trees can also drop their leaves in summer rather than fall in dry years, but usually continue to develop their acorns through the fall. Drought may cause trees not to flower in spring. [5]

Galls on leaf Quercus douglasii-Galls.jpg
Galls on leaf

Gall wasps

Author Ron Russo writes that Q. douglasii hosts the "largest number of known species" of gall wasps, [18] at more than 50. [19] The wasps trigger the formation of oak galls in a wide variety of shapes, colors, and sizes. [18] [19]

Sudden oak death resistance

As of 2002, the disease known as sudden oak death, caused by the oomycete Phytophthora ramorum, had not been found in Quercus douglasii or any members of the white oak group. An experiment showed that Q. douglasii and Q. lobata (another white oak) appeared to be resistant to the pathogen. [13]

Uses

Native Californians commonly gathered Quercus douglasii acorns, which they considered good-tasting, and processed them into acorn flour. They made baskets out of blue oak seedlings, utensils such as bowls from the wood, and dye from the acorns. [5] [20] Commercially, the blue oak is mainly limited to use as firewood. [4] The acorns are eaten by wildlife and livestock. [21] They can be eaten but, if bitter, may need to have the tannins leached. [22]

See also

Notes

  1. Not to be confused with Quercus gambelii Nutt., a separate species.

Related Research Articles

<span class="mw-page-title-main">Oak</span> Tree or shrub in the genus Quercus

An oak is a hardwood tree or shrub in the genus Quercus of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne in a cup. The genus is widely distributed in the Northern Hemisphere; it includes some 500 species, both deciduous and evergreen. Fossil oaks date back to the Middle Eocene. Molecular phylogeny shows that the genus is divided into Old World and New World clades, but many oak species hybridise freely, making the genus's history difficult to resolve.

<i>Quercus kelloggii</i> Species of oak tree

Quercus kelloggii, the California black oak or Kellogg oak, is an oak in the red oak section native to western North America. Although genetically separated from them for more than 20 million years, its leaves are remarkably similar in appearance to several other members of the red oak section including the red oak and the black oak found in eastern and central North America.

<i>Quercus muehlenbergii</i> Species of oak tree

Quercus muehlenbergii, the chinquapinoak, is a deciduous species of tree in the white oak group. The species was often called Quercus acuminata in older literature. Quercus muehlenbergii is native to eastern and central North America. It ranges from Vermont to Minnesota, south to the Florida panhandle, and west to New Mexico in the United States. In Canada it is only found in southern Ontario, and in Mexico it ranges from Coahuila south to Hidalgo.

<i>Quercus garryana</i> Species of oak tree

Quercus garryana is an oak tree species of the Pacific Northwest, with a range stretching from southern California to southwestern British Columbia. It is commonly known as the Garry Oak, Oregon white oak or Oregon oak. It grows from sea level to an altitude of 690 feet in the northern part of its range, and from 980 to 5,900 ft in the south of the range in California. The eponymous Nicholas Garry was deputy governor of the Hudson's Bay Company.

<i>Quercus macrocarpa</i> Species of oak tree

Quercus macrocarpa, the bur oak or burr oak, is a species of oak tree native to eastern North America. It is in the white oak section, Quercus sect. Quercus, and is also called mossycup oak, mossycup white oak, blue oak, or scrub oak. The acorns are the largest of any North American oak, and are important food for wildlife.

<i>Quercus agrifolia</i> Evergreen oak tree native to California

Quercus agrifolia, the California live oak, or coast live oak, is a highly variable, often evergreen oak tree, a type of live oak, native to the California Floristic Province. It may be shrubby, depending on age and growing location, but is generally a medium-sized tree. It grows west of the Sierra Nevada mountain range from Mendocino County, California, south to northern Baja California in Mexico. It is classified in the red oak section of oaks.

<i>Quercus coccinea</i> Species of oak tree

Quercus coccinea, the scarlet oak, is a deciduous tree in the red oak section Lobatae of the genus Quercus, in the family Fagaceae.

<i>Aesculus californica</i> Species of plant

Aesculus californica, commonly known as the California buckeye or California horse-chestnut, is a species of buckeye native to California and southwestern Oregon.

<i>Quercus lobata</i> Species of oak tree

Quercus lobata, commonly called the valley oak or roble, grows into the largest of California oaks. It is endemic to California, growing in interior valleys and foothills from Siskiyou County to San Diego County. Mature specimens may attain an age of up to 600 years. This deciduous oak requires year-round access to groundwater.

<span class="mw-page-title-main">Salinan language</span> Language

Salinan was the indigenous language of the Salinan people of the central coast of California. It has been extinct since the death of the last speaker in 1958.

<span class="mw-page-title-main">California oak woodland</span> Plant community found in the western United States and Mexico

California oak woodland is a plant community found throughout the California chaparral and woodlands ecoregion of California in the United States and northwestern Baja California in Mexico. Oak woodland is widespread at lower elevations in coastal California; in interior valleys of the Coast Ranges, Transverse Ranges and Peninsular Ranges; and in a ring around the California Central Valley grasslands. The dominant trees are oaks, interspersed with other broadleaf and coniferous trees, with an understory of grasses, herbs, geophytes, and California native plants.

<i>Quercus chrysolepis</i> Species of oak tree

Quercus chrysolepis, commonly termed canyon live oak, canyon oak, golden cup oak or maul oak, is a North American species of evergreen oak that is found in Mexico and in the western United States, notably in the California Coast Ranges. This tree is often found near creeks and drainage swales growing in moist cool microhabitats. Its leaves are a glossy dark green on the upper surface with prominent spines; a further identification arises from the leaves of canyon live oak being geometrically flat.

<i>Quercus durata</i> Species of oak tree

Quercus durata, commonly known as leather oak, is a species of oak endemic to California, common in the Coast Ranges and the foothills of the Sierra Nevada. The common name "leather oak" is derived from the leathery texture on the lop of its leaves. Taxonomically it is placed in the white oak group.

<i>Quercus dumosa</i> Species of oak tree

Quercus dumosa is a species of plant in the family Fagaceae, belonging to the white oak section of the oak genus (Quercus). This tree goes by the common names coastal sage scrub oak and Nuttall's scrub oak.

Quercus × alvordiana, the Alvord oak, is a hybrid oak in the genus Quercus. It has been reported to be a hybrid between Quercus douglasii and Quercus turbinella, or between Q. douglasii and Quercus john-tuckeri, which was formerly considered to be a variety of Q. turbinella.

<i>Quercus pagoda</i> Species of oak tree

Quercus pagoda, the cherrybark oak, is one of the most highly valued red oaks in the southern United States. It is larger and better formed than southern red oak and commonly grows on more moist sites. Its strong wood and straight form make it an excellent timber tree. Many wildlife species use its acorns as food, and cherrybark oak makes a fine shade tree. Cherrybark oak was formerly considered to be a subspecies of southern red oak, Quercus falcata, subsp pagodifolia.

<i>Quercus turbinella</i> Species of plant

Quercus turbinella is a North American species of oak known by the common names shruboak, turbinella oak, shrub live oak, and gray oak. It is native to Arizona, California, New Mexico, Utah, Colorado, and Nevada in the western United States. It also occurs in northern Mexico.

<i>Quercus pacifica</i> Species of oak tree

Quercus pacifica is a species of oak known by the common names island scrub oak, Channel Island scrub oak, and Pacific oak.

<i>Notholithocarpus</i> Species of flowering plant in the family Fagaceae

Notholithocarpus densiflorus, commonly known as the tanoak or tanbark-oak, is a broadleaf tree in the family Fagaceae, and the type species of the genus Notholithocarpus. It is native to the far western United States, particularly Oregon and California. It ranges from 15–40 meters in height, with a trunk diameter of 60–190 centimeters.

<span class="mw-page-title-main">Oak woodland</span> Plant community dominated by oaks

An oak woodland is a plant community with a tree canopy dominated by oaks. In terms of canopy closure, oak woodlands are intermediate between oak savanna, which is more open, and oak forest, which is more closed. Although the community is named for the dominance of oak trees, the understory vegetation is often diverse and includes many species of grasses, sedges, forbs, ferns, shrubs, and other plants.

References

  1. Beckman, E. (2016). "Quercus douglasii". IUCN Red List of Threatened Species . 2016: e.T78914533A78914539. doi: 10.2305/IUCN.UK.2016-1.RLTS.T78914533A78914539.en . Retrieved November 12, 2021.
  2. 1 2 Wyly, Zarah (February 8, 2019). "Species Spotlight: Quercus douglasii Hook. & Arn". International Oak Society. Retrieved July 14, 2019.
  3. "Quercus douglasii". World Checklist of Selected Plant Families . Royal Botanic Gardens, Kew via The Plant List. Note that this website has been superseded by World Flora Online
  4. 1 2 3 4 Hogan, C. Michael. "Blue Oak Quercus douglasii". GlobalTwitcher. Archived from the original on May 26, 2013. Retrieved July 14, 2019.
  5. 1 2 3 4 5 6 7 8 9 10 11 12 Fryer, Janet L. (2007). "Quercus douglasii". Fire Effects Information System (FEIS). US Department of Agriculture (USDA), Forest Service (USFS), Rocky Mountain Research Station, Fire Sciences Laboratory.
  6. 1 2 McDonald, Philip M. (1990). "Quercus douglasii". In Burns, Russell M.; Honkala, Barbara H. (eds.). Hardwoods. Silvics of North America. Washington, D.C.: United States Forest Service (USFS), United States Department of Agriculture (USDA). Vol. 2 via Southern Research Station.
  7. "Quercus douglasii". Calflora. Berkeley, California: The Calflora Database.
  8. 1 2 Nixon, Kevin C. (1997). "Quercus douglasii". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 3. New York and Oxford: Oxford University Press via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
  9. Ritter, Lyman V. "Blue Oak Woodland" (PDF). California Department of Fish and Game. Archived from the original (PDF) on August 10, 2009. Retrieved April 12, 2022.
  10. 1 2 "Quercus douglasii Tree Record". SelecTree. Cal Poly San Luis Obispo - Urban Forest Ecosystems Institute. Retrieved July 14, 2019.
  11. Stahle, David. "Ancient Blue Oak Woodlands of California". University of Arkansas Tree-Ring Laboratory. Archived from the original on December 4, 2008. Retrieved July 14, 2019.
  12. "Blue Oak (Quercus douglasii) Species Details and Allergy Info, Santa Clara County, California". PollenLibrary.com.
  13. 1 2 Rizzo, David M.; Garbelotto, Matteo; Davidson, Jennifer M.; Slaughter, Garey W.; Koike, Steven T. (2002). "Phytophthora ramorum and Sudden Oak Death in California: I. Host Relationships" (PDF). U.S. Forest Service. Retrieved July 14, 2019.
  14. Peeters, Hans J.; Peeters, Pam (2005). Raptors of California. University of California Press. p. 147. ISBN   9780520242005.
  15. Tollefson, Jennifer E. (2008). "Quercus chrysolepis". Fire Effects Information System (FEIS). US Department of Agriculture (USDA), Forest Service (USFS), Rocky Mountain Research Station, Fire Sciences Laboratory.
  16. "Plants Profile for Quercus alvordiana (Alvord oak)". Plants Database. USDA Natural Resources Conservation Service. Retrieved July 15, 2019.
  17. Stahle, D. W.; Griffin, R. D.; Meko, D. M.; Therrell, M. D.; Edmondson, J. R.; Cleaveland, M. K.; Stahle, L. N.; Burnette, D. J.; Abatzoglou, J. T.; Redmond, K. T.; Dettinger, M. D.; Cayan, D. R. (May 22, 2013). "The Ancient Blue Oak Woodlands of California: Longevity and Hydroclimatic History". Earth Interactions. 17 (12): 1–23. Bibcode:2013EaInt..17l...1S. doi: 10.1175/2013EI000518.1 . S2CID   130820896.
  18. 1 2 Russo, Ron (July 1, 2009). "Call of the Galls". Bay Nature Magazine. Bay Nature Institure. Retrieved July 15, 2019.
  19. 1 2 Wirka, Jeanne (September 22, 2015). "Nature: Sonoma County is bursting with galls". The Press Democrat. Retrieved July 15, 2019.
  20. Gauna, Forest. "Plant of the Week: Blue Oak (Quercus douglasii Hook. & Arn.)". Celebrating Wildflowers. U.S. Forest Service. Retrieved July 15, 2019.
  21. Little, Elbert L. (1994) [1980]. The Audubon Society Field Guide to North American Trees: Western Region (Chanticleer Press ed.). Knopf. p. 395. ISBN   0394507614.
  22. Elias, Thomas S.; Dykeman, Peter A. (2009) [1982]. Edible Wild Plants: A North American Field Guide to Over 200 Natural Foods. New York: Sterling. pp. 228, 231. ISBN   978-1-4027-6715-9. OCLC   244766414.