The sperm whale or cachalot is the largest of the toothed whales and the largest toothed predator. It is the only living member of the genus Physeter and one of three extant species in the sperm whale family, along with the pygmy sperm whale and dwarf sperm whale of the genus Kogia.
Physeteroidea is a superfamily that includes three extant species of whales: the sperm whale, in the genus Physeter, and the pygmy sperm whale and dwarf sperm whale, in the genus Kogia. In the past, these genera have sometimes been united in a single family, the Physeteridae, with the two Kogia species in the subfamily Kogiinae; however, recent practice is to allocate the genus Kogia to its own family, the Kogiidae, leaving the Physeteridae as a monotypic family, although additional fossil representatives of both families are known.
Thalassocnus is an extinct genus of semiaquatic ground sloths from the Miocene and Pliocene of the Pacific South American coast. It is monotypic within the subfamily Thalassocninae. The five species—T. antiquus, T. natans, T. littoralis, T. carolomartini, and T. yuacensis—represent a chronospecies, a population gradually adapting to marine life in one direct lineage. They are the only known aquatic sloths, but they may have also been adapted to a terrestrial lifestyle. They have been found in the Pisco Formation of Peru, the Tafna Formation of Argentina, and the Bahía Inglesa, Coquimbo, and Horcón formations of Chile. Thalassocninae has been placed in both the families Megatheriidae and Nothrotheriidae.
Zygophyseter varolai is an extinct sperm whale that lived during the Tortonian age of the Late Miocene 11.2 to 7.6 million years ago. It is known from a single specimen from the Pietra Leccese Formation in Italy. It was a member of a stem group of fossil macroraptorial sperm whales also including Brygmophyseter, Acrophyseter, and Livyatan. It probably grew to be around 6.5 to 7 meters in length and shared some characteristics with other raptorials, such as large teeth with tooth enamel that were functional in both the upper and lower jaws which the modern sperm whale lacks. It also had a beak, the ability to echolocate prey, and could have probably swum faster than the modern-day sperm whale which can reach 4 kilometers per hour (2.5 mph). These were probably used in the capture of large prey, such as large fish, seals, and whales. In fact, its common name, the killer sperm whale, refers to its feeding habits that would have had a resemblance to the modern-day killer whale.
Brygmophyseter, known as the biting sperm whale, is an extinct genus of toothed whale in the sperm whale family with one species, B. shigensis. When it was first described in 1994, the species was placed in the genus Scaldicetus based on tooth morphology, but this was later revised in 1995. In 2006, it was classified into the genus Naganocetus, which is considered to be a junior synonym. The only known specimen, a nearly complete skeleton, was dated to be around 16–15 million years old. Brygmophyseter is thought to have been 6.5–7 meters (21–23 ft) long, and it probably had 11 or 12 teeth in the upper and lower jaws. Brygmophyseter is part of a group of macroraptorial sperm whales which tended to be apex predators using their large teeth to catch struggling prey such as whales. It had a spermaceti organ which was probably used for biosonar like in the modern sperm whale. The whale has made an appearance on The History Channel's TV series Jurassic Fight Club.
Livyatan is an extinct genus of macroraptorial sperm whale containing one known species: L. melvillei. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous novel Moby-Dick about a white bull sperm whale. It is mainly known from the Pisco Formation of Peru during the Tortonian stage of the Miocene epoch, about 9.9–8.9 million years ago (mya); however, finds of isolated teeth from other locations such as Chile, Argentina, United States (California), South Africa and Australia imply that either it or a close relative survived into the Pliocene, around 5 mya, and may have had a global presence. It was a member of a group of macroraptorial sperm whales and was probably an apex predator, preying on whales, seals and so forth. Characteristically of raptorial sperm whales, Livyatan had functional, enamel-coated teeth on the upper and lower jaws, as well as several features suitable for hunting large prey.
Orcinus citoniensis is an extinct species of killer whale identified in the Late Pliocene of Italy and the Early Pleistocene of England. It was smaller than the modern killer whale, 4 m (13 ft) versus 7 to 10 m, and had around 8 more teeth in its jaw. It may have resembled the modern killer whale in appearance, and could represent a transitional species between the modern killer whale and other dolphins. O. citoniensis could have hunted fish and squid in pods, and coexisted with other large predators of the time such as the orcinine Hemisyntrachelus and the extinct shark Otodus megalodon.
Diaphorocetus is an extinct genus of odontocete cetacean belonging to Physeteroidea. Its remains were found in the Monte León Formation of Argentina, dating to the Early Miocene.
Scaldicetus is an extinct genus of highly predatory macroraptorial sperm whale. Although widely used for a number of extinct physeterids with primitive dental morphology consisting of enameled teeth, Scaldicetus as generally recognized appears to be a wastebasket taxon filled with more-or-less unrelated primitive sperm whales.
The Pisco Formation is a geologic formation located in Peru, on the southern coastal desert of Ica and Arequipa. The approximately 640 metres (2,100 ft) thick formation was deposited in the Pisco Basin, spanning an age from the Middle Miocene up to the Early Pleistocene, roughly from 15 to 2 Ma. The tuffaceous sandstones, diatomaceous siltstones, conglomerates and dolomites were deposited in a lagoonal to near-shore environment, in bays similar to other Pacific South American formations as the Bahía Inglesa and Coquimbo Formations of Chile.
Albicetus is a genus of stem-sperm whales that lived during the Miocene Epoch, around 15 million years ago, and was discovered in Santa Barbara, California in 1909. It was categorized for decades as belonging to a group of extinct walruses erroneously thought to be sperm whales. It was named Albicetus, meaning "white whale", is a reference to the leviathan in Herman Melville's classic 1851 novel Moby-Dick.
Chilcacetus is an extinct genus of primitive odontocete known from Early Miocene (Aquitanian) of Peru. Fossils were found in and named after the Chilcatay Formation of the Pisco Basin.
Kogia pusilla is an extinct species of sperm whale from the Middle Pliocene of Italy related to the modern day dwarf sperm whale and pygmy sperm whale. It is known from a single skull discovered in 1877, and was considered a species of beaked whale until 1997. The skull shares many characteristics with other sperm whales, and is comparable in size to that of the dwarf sperm whale. Like the modern Kogia, it probably hunted squid in the twilight zone, and frequented continental slopes. The environment it inhabited was likely a calm, nearshore area with a combination sandy and hard-rock seafloor. K. pusilla likely died out due to the ice ages at the end of the Pliocene.
Idiorophus is a genus of toothed whales in the family Physeteridae. Fossils have been found in the Colhuehuapian Gaiman Formation of Argentina and the Libano Sandstone in Italy.
Eudelphis is an extinct genus of sperm whale belonging to Physeteroidea that lived in the ancient North Sea basin about 16-11 million years ago, during the middle Miocene (Langhian).
Macroraptorial sperm whales were highly predatory whales of the sperm whale superfamily (Physeteroidea) of the Miocene epoch that hunted large marine mammals, including other whales, using their large teeth. They consist of five genera: Acrophyseter, Albicetus, Brygmophyseter, Livyatan, and Zygophyseter. All species are known by at least a skull, and are informally grouped without a family designation. They were all likely the apex predator of their habitats, comparable to the modern day killer whale, and achieved great lengths, with one species–Livyatan–measuring about 13.5–17.5 m (44–57 ft).
Hoplocetus is an extinct genus of raptorial cetacean of the sperm whale superfamily, Physeteroidea. Its remains have been found in the Miocene of Belgium, France, Germany and Malta, the Pliocene of Belgium and France, and the Pleistocene of the United Kingdom and South Carolina.
Spheniscus muizoni is an extinct species of banded penguins that lived during the early Late Miocene in what is now Peru, South America. The species, the earliest member of the extant genus, was described in 2007 by Ursula B. Göhlich based on fossils found in the fossiliferous Pisco Formation of the Pisco Basin, southwestern Peru.
Ankylorhiza is an extinct genus of toothed whale that lived in what is now the United States during the Oligocene epoch, between 29 and 23.5 million years ago. The type and only known species is A. tiedemani, though two fossil skeletons may represent an additional, second species within the genus. Ankylorhiza was about 4.8 meters (16 ft) long, with a long, robust skull bearing conical teeth that were angled forwards at the tip of the snout.
