Arandaspida Temporal range: Early Ordovician – Late Ordovician, | |
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Life restoration of Arandaspis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Infraphylum: | Agnatha |
Class: | † Pteraspidomorphi |
Subclass: | † Arandaspida Ritchie and Gilbert-Tomlinson, 1977 |
Families | |
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Arandaspida is a taxon of very early, jawless prehistoric fish which lived during the Ordovician period. Arandaspids represent some of the oldest known vertebrates. The group represents a subclass within the class Pteraspidomorphi, and contains only one order, the Arandaspidiformes. The oldest known genus of this group is Sacabambaspis found in South America.
The head armor of arandaspids is elongated, fusiform, with a rather flat dorsal shield, and a bulging ventral shield. In the anterior part of the dorsal shield are two closely set holes, which have been thought to be a paired pineal opening, but which are more likely the external openings of the endolymphatic ducts.
The eyes, surrounded by a sclerotic ring, are housed in a notch at the anterior end of the dorsal shield. The nostrils are not clearly located, but may have been situated between the eyes. Ventrally, the ventral lip of the mouth is armed with long series of small oral plates which recall those of heterostracans.
The gill openings are probably numerous (more than 15) and minute. They opened between the diamond-shaped platelets which separate the dorsal from the ventral shield.
The body is covered with rod-shaped scales arranged in chevrons, and the tail is probably pad-shaped and diphycercal. The dermal bones of arandaspids consist of aspidine (acellular bone) and are ornamented with oakleaf-shaped tubercles which seem to contain no dentine. The sensory-lines were housed in narrow grooves between the tubercles.
Taxonomy based on the work of Mikko's Phylogeny Archive, [2] Nelson, Grande and Wilson 2016 [3] and van der Laan 2018. [4]
Heterostraci is an extinct subclass of pteraspidomorph, Ostracoderm, jawless vertebrate that lived primarily in marine and estuary environments. Heterostraci existed from the mid-Ordovician to the conclusion of the Devonian.
Lepidaspis serrata is an extinct heterostracan jawless fish from Early Devonian Canada. Its scientific name refers to the fact that the armor is composed of hundreds of tiny scales with serrated edges.
Anaspida is an extinct group of jawless fish that existed from the early Silurian period to the late Devonian period. They were classically regarded as the ancestors of lampreys, but it is denied in recent phylogenetic analysis, although some analysis show these group would be at least related. Anaspids were small marine fish that lacked a heavy bony shield and paired fins, but were distinctively hypocercal.
Pteraspidomorphi is an extinct class of early jawless fish. They have long been regarded as closely related or even ancestral to jawed vertebrates, but the few characteristics they share with the latter are now considered as basal traits for all vertebrates.
Eriptychiida is an extinct marine taxon of vertebrate in the group Pteraspidomorphi.
This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes.
Arandaspis prionotolepis is an extinct species of jawless fish that lived in the Ordovician period, about 480 to 470 million years ago. Its remains were found in the Stairway Sandstone near Alice Springs, Australia in 1959, but it was not determined that they were the oldest known vertebrates until the late 1960s. Arandaspis is named after a local Indigenous Australian people, the Aranda.
Drepanaspis is an extinct genus of heterostracan armoured jawless fish from the Early Devonian. Drepanaspis are assumed to have lived primarily in marine environments and is most commonly characterized by their ray-like, heavily armoured bodies, along with their lack of paired fins and jaws.
Groenlandaspis is an extinct genus of arthrodire from the Late Devonian. Fossils of the different species are found in late Devonian strata in all continents except eastern Asia. The generic name commemorates the fact that the first specimens of the type species were found in Greenland.
Astraspis is an extinct, monotypic genus of primitive jawless fish from the Ordovician of Central North America including the Harding Sandstone of Colorado and Bighorn Mountains of Wyoming. It is also known from Bolivia. It is related to other Ordovician fishes, such as the South American Sacabambaspis, and the Australian Arandaspis.
Micropogonias is a genus of marine ray-finned fishes belonging to the family Sciaenidae, the drums and croakers. These fishes are found in the eastern Pacific and western Atlantic Oceans.
Astraspida, or astraspids, are a small group of extinct armored jawless vertebrates, which lived in the Late Ordovician in North America. They are placed among the Pteraspidomorphi because of the large dorsal and ventral shield of their head armor. They are represented by a single genus, Astraspis, including possibly two species, A. desiderata and A. splendens but their remains are fairly abundant in Ordovician sandstones of the USA and Canada (Quebec). The head armor of Astraspis is rather massive, with a series of ten gill openings lining the margin of the dorsal shield, and laterally placed eyes. The dorsal shield is ribbed by strong longitudinal crests, and the tail is covered with large, diamond-shaped scales. They are often grouped together with the Arandaspidida.
Sacabambaspis is an extinct genus of jawless fish that lived in the Ordovician period. Sacabambaspis lived in shallow waters on the continental margins of Gondwana. It is the best known arandaspid with many specimens known.
Eusthenodon is an extinct genus of tristichopterid tetrapodomorphs from the Late Devonian period, ranging between 383 and 359 million years ago. They are well known for being a cosmopolitan genus with remains being recovered from East Greenland, Australia, Central Russia, South Africa, Pennsylvania, and Belgium. Compared to the other closely related genera of the Tristichopteridae clade, Eusthenodon was one of the largest lobe-finned fishes and among the most derived tristichopterids alongside its close relatives Cabonnichthys and Mandageria.
Ogcocephalus is a genus of marine ray-finned fishes belonging to the family Ogcocephalidae, the deep sea batfishes. The species in this genus are found in the Eastern Pacific Ocean and Western Atlantic Ocean, with most species in the eastern Atlantic Ocean where they live at shallower depths than the other ogcocephalid genera.
Ptycholepiformes are an extinct order of prehistoric ray-finned fish that existed during the Triassic period and the Early Jurassic epoch. The order includes the genera Acrorhabdus, Ardoreosomus, Boreosomus, Chungkingichthys, Ptycholepis, and Yuchoulepis. Although several families have been proposed, some studies place all these genera in the same family, Ptycholepididae.
The deceitful velvetfish is a species of marine ray-finned fish, a velvetfish, belonging to the family Aploactinidae. This species is endemic to the oceans around Australia. This species is the only known member of its genus.
Cipactlichthys is a genus of extinct marine holostean fish from the Lower Cretaceous of Mexico, distantly related to modern bowfins. The only known species is Cipactlichthys scutatus.
Microbrachius is an extinct genus of tiny, advanced antiarch placoderms closely related to the bothriolepids. Specimens range in age from the Lower Devonian Late Emsian Stage to the Middle Devonian Upper Givetian Stage. They are characterized by having large heads with short thoracic armor of an average length of 2–4 cm. There are patterns of small, but noticeable tubercles on the armor, with the arrangement varying from species to species. Specimens of Microbrachius have been found in Scotland, Belarus, Estonia, and China.
Asterolepis is an extinct genus of antiarch placoderms from the Devonian of North and South America and Europe. They were heavily armored flat-headed benthic detritivores with distinctive jointed limb-like pectoral fins and hollow spine. The armor plate gives the Asterolepis a box-like shape. Its pectoral fins are also armored but the caudal and dorsal fin are not. The first fossils were named by M. Eichwald in 1840 after noticing star-like markings on the fossils.
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