Arhopala | |
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Arhopala japonica is of unclear affiliations. When Narathura was considered distinct, it was placed there. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Lepidoptera |
Family: | Lycaenidae |
Tribe: | Arhopalini |
Genus: | Arhopala Boisduval, 1832 |
Type species | |
Arhopala phryxus Boisduval, 1832 | |
Diversity | |
About 220 species | |
Synonyms | |
Several, see text |
Arhopala is a very large genus of gossamer-winged butterflies (Lycaenidae). They are the type genus of the tribe Arhopalini. In the relatively wide circumscription used here, it contains over 200 species collectively known as oakblues. They occur from Japan throughout temperate to tropical Asia south and east of the Himalayas to Australia and the Solomon Islands of Melanesia. Like many of their relatives, their caterpillars are attended and protected by ants (myrmecophily). Sexual dichromatism is often prominent in adult oakblues. [1] [ failed verification ]
The genus' delimitation versus Amblypodia and Flos has proven to be problematic; not all issues are resolved and the assignment of species to these genera must be considered somewhat provisional.
As circumscribed here, this genus includes many formerly independent genera. Junior synonyms of Arhopala are: [1]
The species have been provisionally sorted into groups of presumed closest relatives, but many species remain insufficiently studied even for such a preliminary assessment at present. For example, A. phryxus – the type species used by Jean Baptiste Boisduval when he described Arhopala in 1832 – was established at the same time as and specifically for this genus. It is considered to be a valid species of unclear affiliations, but it is suspected that Boisduval's taxon is a junior synonym of A. thamyras – the namesake of its species group – which had been described as Papilio thamyras by Carl Linnaeus already in 1764. [2]
Molecular phylogenetic studies have only sampled a fraction of the known diversity of oakblues, but as it seems at least some of the groups represent clades that could justifiably be treated as subgenera. It is also to be seen, however, if the genus is monophyletic in the loose sense as used here, or would need to be split up again. [1]
Species are listed alphabetically, while the groups are listed in the presumed phylogenetic sequence: [2]
centaurus group (subgenus Nilasera if valid)
agesias group
anthelus group (subgenus Narathura if valid)
camdeo group
oenea group
epimuta group
amphimuta group
agesilaus group
alitaeus group
| agrata group
cleander group
eumolphus group
rama group
agelastus group
fulla group
aurea group (subgenus Aurea if valid)
ganesa group (subgenus Panchala if valid)
abseus group
thamyras group
hercules group
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The Heliconiinae, commonly called heliconians or longwings, are a subfamily of the brush-footed butterflies. They can be divided into 45–50 genera and were sometimes treated as a separate family Heliconiidae within the Papilionoidea. The colouration is predominantly reddish and black, and though of varying wing shape, the forewings are always elongated tipwards, hence the common name.
The Limenitidinae are a subfamily of butterflies that includes the admirals and relatives. The common names of many species and genera reference military ranks or – namely the Adoliadini – titles of nobility, in reference to these butterflies' large size, bold patterns, and dashing flight. In particular, the light stripe running lengthwise across the wings of many Limenitidini has reminded earlier authors of officers' shoulder marks and epaulets.
The Pyralini are a tribe of snout moths described by Pierre André Latreille in 1809. They belong to the subfamily Pyralinae, which contains the "typical" snout moths of the Old World and some other regions. The genus list presented here is provisional.
The Eumaeini are a tribe of gossamer-winged butterflies. They are typically placed in the subfamily Theclinae, but sometimes considered a separate subfamily Eumaeinae. Over 1,000 species are found in the Neotropical realm
Arhopala alax, the silky oakblue, is a small butterfly found in India that belongs to the lycaenids or blues family.
Lamasina is a genus of gossamer-winged butterflies ; the validity of its name is subject to dispute. Among its family, these sexually dimorphic Lepidoptera belong to the tribe Eumaeini of the subfamily Theclinae. Lamasina species are found mainly in northern South America, approximately to the Guyanas. L. draudti is also found in Central America south of the Yucatán Peninsula. In the Andes, the genus extends somewhat further south; L. rhaptissima almost reaches Bolivia.
Phengaris is a genus of gossamer-winged butterflies in the subfamily Polyommatinae. Commonly, these butterflies are called large blues, which if referring to a particular species is P. arion, a species resident in Europe and some parts of Asia.
Arhopala aida or white-stained oakblue, is a species of butterfly belonging to the lycaenid family described by Lionel de Nicéville in 1889. It is found in Southeast Asia . The upper surface of the male is lilac the black marginal band on both wings above equally broad. The under surface is brown with a distinct violet gloss;forewing with a light hindmarginal area. The female is above bluer, lighter than the male, and has a much broader black margin. aida differs from Ahropalus myrtale in its darker and more violet-blue and its broad outer margins ; the undersides are, however, very much alike.
Arhopala athada, the vinous oakblue is a species of butterfly belonging to the lycaenid family described by Otto Staudinger in 1889. It is found in Southeast Asia - Singapore, Peninsular Malaya, Sumatra, Borneo, Bangka, Bawean, Assam, Burma, Mergui, Thailand and the Philippines.
Arhopala lurida is a species of butterfly belonging to the lycaenid family. It is found in Southeast Asia
Arhopala muta is a species of butterfly belonging to the lycaenid family described by William Chapman Hewitson in 1862. It is found in Southeast Asia
Arhopala perimuta, the yellowdisc oakblue or yellowdisc tailless oakblue, is a species of butterfly belonging to the lycaenid family described by Frederic Moore in 1857. It is found in Southeast Asia.
Arhopala pseudomuta, Raffles's oakblue, is a species of butterfly belonging to the lycaenid family. It was described by Otto Staudinger in 1889. It is found in Southeast Asia - Peninsular Malaya, Singapore pseudomuta),Langkawi, Mergui, Burma, Thailand ,Sumatra, Borneo
Arhopala vihara,the Large Spotted Oakblue, is a species of butterfly belonging to the lycaenid family described by (Cajetan Felder and Rudolf Felder in 1860. It is found in Southeast Asia.
Arhopala zambra, the Zambra oakblue, is a species of butterfly belonging to the lycaenid family described by Charles Swinhoe in 1911. It is found in Southeast Asia.
Arhopala camdeo, the lilac oakblue, is a butterfly in the family Lycaenidae. It was described by Frederic Moore in 1858. It is found in the Indomalayan realm.
Arhopala thamyras is a butterfly in the family Lycaenidae. It was described by Carl Linnaeus in 1758. It is found in the Australasian realm.
Arhopala khamti, Doherty's dull oakblue, is a butterfly in the family Lycaenidae. It was described by William Doherty in 1891. It is found in the Indomalayan realm.
Arhopala moolaiana or pale yellow oakblue, is a butterfly in the family Lycaenidae. It was described by Frederic Moore in 1879. It is found in the Indomalayan realm.
Arhopala asopia, the plain tailless oakblue, is a butterfly in the family Lycaenidae. It was described by William Chapman Hewitson in 1869. It is found in the Indomalayan realm.