Hortalotarsus Temporal range: Early Jurassic, | |
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Preserved portion of the holotype drawn before being partially destroyed by blasting | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Bagualosauria |
Clade: | † Plateosauria |
Clade: | † Massopoda |
Family: | † Massospondylidae |
Genus: | † Hortalotarsus Seeley, 1894 |
Type species | |
†Hortalotarsus skirtopodus Seeley, 1894 | |
Synonyms | |
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Hortalotarsus (etymology uncertain; probably "tarsus of a young bird"? [1] ) is a dubious genus of extinct sauropodomorph from the Early Jurassic (Sinemurian-aged) Clarens Formation of Eagle's Crag, South Africa. [2] [3]
The type species, Hortalotarsus skirtopodus was named by Harry Seeley in 1894, initially as a species of Thecodontosaurus . [4] The holotype is AM 455, [5] consisting today of only a tibia, fibula and phalanges, although more of the skeleton was initially present, including ribs, possible vertebrae, a possible ilium, a possible femur, a possible scapula and a possible humerus, [4] [6] which was discovered by William Horner Wallace on 11 July 1888 in Eagle's Crag, Barkly East. [7]
According to Robert Broom (1911), "Originally most of the skeleton was in the rock, and it was regarded by the farmers as the skeleton of a Bushman, but it is said to have been destroyed through fear that a Bushman skeleton in the rock might tend to weaken the religious belief of the rising generation." [6] Seeley however, states that most of the skeleton was lost by a failed attempt to free it from the rock by using gunpowder. [4] Some partial leg bones were salvaged. [8]
Hortalotarsus would have been similar to Massospondylus, [8] reaching around 3 metres (9.8 ft) long when fully grown.
Hortalotarsus was subsequently regarded as either a synonym of Massospondylus [5] or a valid genus belonging to Anchisauridae. [9] [10] Galton and Cluver (1976) as well as Galton and Upchurch (2004), however, designated this genus a nomen dubium. [8] [11]
Today, Hortalotarsus is classified as a member of the Massospondylidae. [8]
Sauropodomorpha is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The prosauropods, which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic until their decline and extinction at the end of the Cretaceous.
Anchisaurus is a genus of basal sauropodomorph dinosaur. It lived during the Early Jurassic Period, and its fossils have been found in the red sandstone of the Upper Portland Formation, Northeastern United States, which was deposited from the Hettangian age into the Sinemurian age, between about 200 and 192 million years ago. Until recently it was classed as a member of Prosauropoda. The genus name Anchisaurus comes from the Greek αγχιanchi-; "near, close" + Greek σαυρος ; "lizard". Anchisaurus was coined as a replacement name for "Amphisaurus", which was itself a replacement name for Hitchcock's "Megadactylus", both of which had already been used for other animals.
Thecodontosaurus is a genus of herbivorous basal sauropodomorph dinosaur that lived during the late Triassic period.
Massospondylus was a genus of sauropodomorph dinosaur from the Early Jurassic. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been named. Fossils have since been found at other locations in South Africa, Lesotho, and Zimbabwe. Material from Arizona's Kayenta Formation, India, and Argentina has been assigned to the genus at various times, but the Arizonan and Argentinian material are now assigned to other genera.
Lufengosaurus is a genus of massospondylid dinosaur which lived during the Early Jurassic period in what is now southwestern China.
Euskelosaurus is a sauropodomorph dinosaur from the Late Triassic of South Africa and Lesotho. Fossils have only been recovered from the lower Elliot Formation in South Africa and Lesotho, and in one locality in Zimbabwe.
Coloradisaurus is a genus of massospondylid sauropodomorph dinosaur. It lived during the Late Triassic period in what is now La Rioja Province, Argentina. It is known from two specimens collected from the Los Colorados Formation of the Ischigualasto-Villa Unión Basin.
Gigantoscelus is a dubious genus of basal sauropodomorph dinosaur from the Early Jurassic of South Africa.
Gryponyx is an extinct genus of massopod sauropodomorph known from southern Free State, central South Africa.
Gyposaurus is a genus of basal sauropodomorph dinosaur from the early Jurassic of South Africa. It is usually considered to represent juveniles of other prosauropods, but "G." sinensis is regarded as a possibly valid species.
Plateosauridae is a family of plateosaurian sauropodomorphs from the Late Triassic of Europe, Greenland, Africa and Asia. Although several dinosaurs have been classified as plateosaurids over the years, the family Plateosauridae is now restricted to Plateosaurus, Yimenosaurus, Euskelosaurus, and Issi. In another study, Yates (2003) sunk Sellosaurus into Plateosaurus. Gresslyosaurus is alternatively considered its own genus or a synonym of Plateosaurus.
Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and Antarctica during the Late Triassic to the Early Jurassic periods. Several dinosaurs have been classified as massospondylids over the years. The largest cladistic analysis of early sauropodomorphs, which was presented by Apaldetti and colleagues in November 2011, found Adeopapposaurus, Coloradisaurus, Glacialisaurus, Massospondylus, Leyesaurus and Lufengosaurus to be massospondylids. This result supports many previous analyses that tested fewer taxa. However, this analysis found the two recently described North American massopods, Sarahsaurus and Seitaad, and the South African Ignavusaurus to nest outside Massospondylidae, as opposed to some provisional proposals. Earlier in 2011, Pradhania, a sauropodomorph from India, was tested for the first time in a large cladistic analysis and was found to be a relatively basal massospondylid. Mussaurus and Xixiposaurus may also be included within Massospondylidae.
Pradhania is a genus of massospondylid sauropodomorph dinosaur from the Sinemurian-age Upper Dharmaram Formation of India. It was first named by T. S. Kutty, Sankar Chatterjee, Peter M. Galton and Paul Upchurch in 2007 and the type species is Pradhania gracilis. It was a sauropodomorph of modest size, only about four meters (13 ft) long, and is known from fragmentary remains. It was originally regarded as a basal sauropodomorph but new cladistic analysis performed by Novas et al., 2011 suggests that Pradhania is a massospondylid. Pradhania presents two synapomorphies of Massospondylidae recovered in their phylogenetic analysis.
Lamplughsaura is a genus of sauropodomorph dinosaur from the Sinemurian-age Dharmaram Formation of India, dating from between 196 and 190 million years ago. The type and only species is Lamplughsaura dharmaramensis. It is known from several partial skeletons of a large quadrupedal animal up to 10 meters (33 ft) long, and was either a basal sauropod or, less likely, a more basal sauropodomorph.
Anchisauria is an extinct clade of sauropodomorph dinosaurs that lived from the Late Triassic to the Late Cretaceous. The name Anchisauria was first used Haekel and defined by Galton and Upchurch in the second edition of The Dinosauria. It is a node-based taxon containing the most recent common ancestor of Anchisaurus polyzelus and Melanorosaurus readi, and all its descendants. Galton and Upchurch assigned a family of dinosaurs to the Anchisauria: the Melanorosauridae. The more common prosauropods Plateosaurus and Massospondylus were placed in the sister clade Plateosauria.
The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, southern Free State, and in the eastern KwaZulu-Natal provinces of South Africa. Outcrops and exposures are also found in several localities in Lesotho such as Qacha's Neck, Hill Top, Quthing, and near the capital, Maseru. The Elliot Formation is further divided into the lower (LEF) and upper (UEF) Elliot formations to differentiate significant sedimentological differences between these layers. The LEF is dominantly Late Triassic (Norian-Hettangian) in age while the UEF is mainly Early Jurassic (Sinemurian-Pliensbachian) and is tentatively regarded to preserve a continental record of the Triassic-Jurassic boundary in southern Africa. This geological formation is named after the town of Elliot in the Eastern Cape, and its stratotype locality is located on the Barkly Pass, 9 km north of the town.
Adeopapposaurus is a genus of plateosaurian dinosaur from the Early Jurassic Cañón del Colorado Formation of San Juan, Argentina. It was similar to Massospondylus. Four partial skeletons with two partial skulls are known.
Phytodinosauria is a group of dinosaurs proposed in 1986, combining the Sauropodomorpha and Ornithischia as sister groups, conceptualized as a superorder of herbivorous dinosaurs excluding the carnivorous Theropoda. This hypothesis has been refuted by modern cladistic analysis, showing such a group to be polyphyletic. Modern studies either combine the Theropoda and Sauropodormorpha in the Saurischia or the Theropoda and Ornithischia in the Ornithoscelida.
Ignavusaurus is a genus of basal sauropodomorph dinosaur that lived during the Early Jurassic in what is now Lesotho. Its fossils were found in the Upper Elliot Formation which is probably Hettangian in age. It was described on the basis of a partial, well preserved articulated skeleton. The type species, I. rachelis, was described in 2010 by Spanish palaeontologist F. Knoll.
Ngwevu is a genus of massospondylid sauropodomorph dinosaur from the Lower Jurassic of South Africa. The genus contains one species, Ngwevu intloko.