| Mayaheros urophthalmus | |
|---|---|
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Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| Order: | Cichliformes |
| Family: | Cichlidae |
| Genus: | Mayaheros |
| Species: | M. urophthalmus |
| Binomial name | |
| Mayaheros urophthalmus | |
| Synonyms | |
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Mayaheros urophthalmus, also known as the Mayan cichlid or Mexican mojarra is a species of cichlid.
Previously, this species was a member of the genus Cichlasoma and the subgeneric section Nandopsis . However, the genus Cichlasoma is in revision and has been restricted to the 12 species of South American cichlids related to Cichlasoma bimaculatum. Thus, many of the approximately 100 species that were formerly members of Cichlasoma have yet to be formally assigned to new genera. [2]
This species can be found in Middle America. It is native to the Atlantic slope of tropical Mesoamerica, ranging from eastern Mexico southward to Nicaragua. [3] It was first recorded from Everglades National Park, Florida in 1983 and is now a common nonindigenous fish in South Florida. [4]
The Mayan cichlid inhabits freshwater marshes, mangrove swamps, lakes, rivers, rocky shorelines, lagoons, estuaries, and coastal islands. Adults prefer coastal lagoons and rivers and may survive in marine conditions. [5] It can be found in oxygen-rich areas near submerged vegetation and over muddy substrates. Despite its preference for waters with dissolved oxygen content of at least 3.5 mg/L, though, it is capable of surviving in extreme hypoxia, because it is an oxygen conformer, becoming much less active in hypoxic water, and even surviving virtual anoxia for up to two hours. [6] Unusually, this fish has been recorded in some cenotes in the Yucatan Peninsula.
Mayaheros urophthalmus reaches a maximum length of 39.4 cm (15.5 in). [5] This fish is one of the larger cichlids. The body is oval and flattened on the sides and the head is tapering towards the mouth. Fins are spiny. It has a base color of yellow-brown to gray-brown in most cases, that becomes intense red during breeding. The head and throat are reddish, especially in younger fish. The caudal fin and the soft-sections of the dorsal and anal fin are usually more or less reddish. Six wide green-black vertical stripes run across the sides of the body between the base of the pectoral fin and the base of the caudal fin. There is a large black eye spot circled with blue-green on the tail stem (hence the species scientific name urophthalmus (from oura, Greek for tail and ophthalmos, Greek for eye). [7] Since the distribution area of the species is relatively large, this species may occurs in different colors, which may differ from the one described above.
In its native range, the Mayan cichlid is a popular food fish. For this reason it is the basis of a regional fishery, is commonly used in aquaculture. As in many animals, the red color is much more brilliant in wild specimens than captive ones, but one can help maintain some of its vibrance by feeding the fish live foods and foods that contain Vitamin A, which breaks down into the red pigment beta-carotene in the body.
It is among the most-studied of the Neotropical cichlids. [8] To date, the Mayan cichlid has been most intensively studied at localities in southeastern Mexico on or near the Yucatán Peninsula. [6]
In Thailand, it is listed as one of 13 aquatic species prohibited for import and possession including cultivation by the Department of Fisheries (DOF) alongside other species such as zebra tilapia (Heterotilapia buttikoferi) and blackchin tilapia (Sarotherodon melanotheron) etc. [9]
The Mayan cichlid is philopatric, or site tenacious, i.e. - individuals are non-migratory and prefer to stay within a home range. [10] The Mayan cichlid has a minimum temperature requirement of about 14 °C (57 °F). [11] In its native range, it inhabits waters with temperatures from 18–34 °C (64–93 °F), but its optimal temperature range is 28–33 °C (82–91 °F). [12]
The Mayan cichlid is capable of surviving in a variety of conditions. It is euryhaline and can survive in a range of salinity from 0 – 40 ppt. [8] Experiments on captive specimens have shown that it can tolerate abrupt increases in salinity of up to 15 ppt. [13] The Mayan cichlid has a broad range of tolerance to abiotic conditions and a broad functional repertoire to enable it to feed on about 20% of evasive prey due to its 6.8% jaw protrusion while feeding. [14]
This species is also a dietary generalist, consuming organisms from a variety of disparate taxa. [15] However, it is susceptible to malnourishment, apparently due to the requirement that a relatively large proportion of its diet be animal prey. [8] The Mayan cichlid's diet includes plant matter, smaller fish, algae, detritus, snails, crustaceans, and insects. [4] [16]
The Mayan cichlid is territorial and aggressive when breeding. [5] As parents, Mayan cichlids are highly protective of their young, and they have several broods per year. This species is a monogamous, biparental substrate spawner that exhibits minimal sexual dimorphism and guards its fry for up to six weeks. All of these traits are greatly developed and represent an extreme in the general pattern found in the genus Mayaheros . In Mexico, the Mayan cichlid spawns for a nine-month period from March to November, particularly during the wet season from June to September. This corresponds to a period when the water temperature is at least 24 degrees Celsius. [8] Multiple broods are raised per year. The fry appear to be adapted to lotic water. They exhibit strongly positive geotactic behavior, actively swimming down to the substrate upon hatching from the egg and adhering themselves to the bottom by means of three pairs of mucous glands. [8]
Cichlids are fish from the family Cichlidae in the order Cichliformes. Traditionally Cichlids were classed in a suborder, the Labroidei, along with the wrasses (Labridae), in the order Perciformes, but molecular studies have contradicted this grouping. On the basis of fossil evidence, it first appeared in Argentina during the Early Eocene epoch, about 48.6 million years ago; however, molecular clock estimates have placed the family's origin as far back as 67 million years ago, during the late Cretaceous period. The closest living relative of cichlids is probably the convict blenny, and both families are classified in the 5th edition of Fishes of the World as the two families in the Cichliformes, part of the subseries Ovalentaria. This family is large, diverse, and widely dispersed. At least 1,650 species have been scientifically described, making it one of the largest vertebrate families. New species are discovered annually, and many species remain undescribed. The actual number of species is therefore unknown, with estimates varying between 2,000 and 3,000.
Tilapia is the common name for nearly a hundred species of cichlid fish from the coelotilapine, coptodonine, heterotilapine, oreochromine, pelmatolapiine, and tilapiine tribes, with the economically most important species placed in the Coptodonini and Oreochromini. Tilapia are mainly freshwater fish inhabiting shallow streams, ponds, rivers, and lakes, and less commonly found living in brackish water. Historically, they have been of major importance in artisanal fishing in Africa, and they are of increasing importance in aquaculture and aquaponics. Tilapia can become a problematic invasive species in new warm-water habitats such as Australia, whether deliberately or accidentally introduced, but generally not in temperate climates due to their inability to survive in cold water.
Cichlasoma is a genus of freshwater fish in the cichlid family. The genus was previously very large, including cichlids from North America, including Central America, and South America.
Hemichromis is a genus of fishes from the cichlid family, known in the aquarium trade as jewel cichlids. Jewel cichlids are native to Africa. Within West Africa, Hemichromis species are found in creeks, streams, rivers and lakes with a variety of water qualities including brackish water lagoons. As traditionally defined, the genus includes two distinctly different groups: The five-spotted cichlid group and the true jewel cichlid group (Rubricatochromis), which sometimes are recognized as distinct genera.
Peacock bass or Brazilian tucunaré are large freshwater cichlids of the genus Cichla. These are diurnal predatory fishes native to the Amazon and Orinoco basins, as well as rivers of the Guianas, in tropical South America. They are sometimes referred to in English by their Brazilian name tucunaré or their Spanish name pavon. Despite the common name and their superficial similarity, they are not closely related to other fish known as bass, such as the North American largemouth bass.
The convict cichlid is a fish species from the family Cichlidae, native to Central America, also known as the zebra cichlid. Convict cichlids are popular aquarium fish and have also been the subject of numerous studies on fish behaviour.
Oreochromis is a large genus of oreochromine cichlids, fishes endemic to Africa and the Middle East. A few species from this genus have been introduced far outside their native range and are important in aquaculture. Many others have very small ranges; some are seriously threatened, and O. ismailiaensis and O. lidole possibly are extinct. Although Oreochromis primarily are freshwater fish of rivers, lakes and similar habitats, several species can also thrive in brackish waters and some even survive in hypersaline conditions with a salinity that far surpasses that of seawater. In addition to overfishing and habitat loss, some of the more localized species are threatened by the introduction of other, more widespread Oreochromis species into their ranges. This is because they—in addition to competing for the local resources—often are able to hybridize.
Vieja is a genus of cichlid fish from Central America and Mexico. The majority of the species are freshwater fish found in stagnant or slow-moving waters of southern Mexico to El Salvador, but V. maculicauda, which also occurs in brackish waters, ranges south to Panama. They are high-bodied cichlids that reach lengths of up to 17–35 cm (7–14 in) depending on the exact species. Vieja feed mostly on vegetable matter, but may also take small invertebrates.
Amphilophus is a genus of cichlid fishes from Central America, ranging from southern Mexico to Panama. The genus currently contains 23 species, including several that are well known from the aquarium trade. However, studies led by Oldřich Říčan in 2008 and 2016 suggested that several species within Amphilophus should be moved to the genus Astatheros. Species proposed to be moved to Astatheros in 2008 were A. alfari, A. altifrons, A. bussingi, A. diquis, A. longimanus, A. macracanthus, A. margaritifer, A. rhytisma, A. robertsoni and A. rostratus. Further genetic studies led Říčan to put A. macracanthus in Astatheros, but to put A. alfari, A. altifrons, A. bussingi, A. diquis, A. longimanus, A. rhytisma, A. robertsoni and A. rostratus within the genus Cribroheros. Říčan's study suggests that the Astatheros species are more closely related to the Jack Dempsey and rainbow cichlid than to the remaining Amphilophus species.
A brackish-water aquarium is an aquarium where the water is brackish (semi-salty). The range of "saltiness" varies greatly, from near freshwater to near marine and is often referred to as specific gravity (SG) or salinity. Brackish water aquaria is a popular specialization within the fishkeeping hobby. Many species of fish traded as freshwater species are actually true brackish species, for example mollies, Florida flagfish, and some cichlids such as chromides and black-chin tilapia. There are also several popular species traded purely as brackish water fish, including monos, scats, archerfish, and various species of pufferfish, goby, flatfish, and gar. Generally, aquarists need to maintain a specific gravity of around 1.005 to 1.010 depending on the species being kept, but practically all brackish water fish tolerate variations in salinity well, and some aquarists maintain that regularly fluctuating the salinity in the aquarium actually keeps the fish healthy and free of parasites.
Herotilapia multispinosa also known as the rainbow cichlid is a Central American freshwater fish of the cichlid family. It is found on the Atlantic slope of Honduras, Nicaragua, and Costa Rica from Patuca River (Honduras) south to Matina River, and on the Pacific slope of Nicaragua and Costa Rica from Guasaule River south to Tempisque River. Specimens are also reported from the Choluteca River on the Pacific side of Honduras. This species is found in lakes and swamps with muddy bottoms, where it uses its specialized teeth and only 3.5% jaw protrusion to feed mostly on algae. It is commercially important as an aquarium fish. The rainbow cichlid prefers a pH range of 7.0–8.0, water hardness of 9-20 dGH and a temperature range of 21–36 °C.
Throughout much of the tropics, tilapiine cichlids native to Africa and the Levant have been widely introduced into a variety of aquatic systems. In the U.S. states of Florida and Texas, tilapia were originally introduced to curtail invasive plants. In an effort to meet the growing demand for tilapia, humans have farmed these fish in countries around the world. Capable of establishing themselves into new ponds and waterways, many tilapia have escaped aquaculture facilities across much of Asia, Africa, and South America. In other cases, tilapia have been established into new aquatic habitats via aquarists or ornamental fish farmers.
Alcolapia grahami, the Lake Magadi tilapia or Graham's cichlid, is a vulnerable species of fish in the family Cichlidae. It is specialised to live in hot, alkaline waters in springs and lagoons around hypersaline lakes.
The Salvin's cichlid, also known as the yellow-belly cichlid or tricolored cichlid, is a species of the family Cichlidae. It is found in rivers of the Atlantic slope of southern Mexico, Belize, and Guatemala. It is the only species in the genus Trichromis.
The Cichlasomatinae are a subfamily of cichlid fishes, including all cichlids native to the Greater Antilles, United States, Mexico and Central America, and many of the cichlids from South America. The subfamily Cichlasomatinae is often divided into two tribes: Cichlasomatini and Heroini, however some authorities classify these two tribes as part of the wider Neotropical and marginally Nearctic subfamily Cichlinae.
Rocio is a small genus of cichlid freshwater fishes from southern Mexico and northern Central America.
The three spot cichlid, also known as the trimac or red-eyed cichlid, is a species of cichlid from Mexico and Central America, from the subfamily Cichlasomatinae. It is rarely found as an aquarium fish. Although still included as a highly aberrant member of Cichlasoma by FishBase, other authorities such as Catalog of Fishes have moved it to the genus Amphilophus.
Heroini is a fish tribe from the Cichlasomatinae subfamily in the family Cichlidae (cichlids). All cichlids native to the Greater Antilles, United States, Mexico and northern Central America are members of this tribe. It also includes most cichlid species in southern Central America and several species from South America. A large percentage of its species were formerly placed in the genus Cichlasoma but have since been moved to other genera.
The redbelly tilapia, also known as the Zille's redbreast tilapia or St. Peter's fish, is a species of fish in the cichlid family. This fish is found widely in fresh and brackish waters in the northern half of Africa and the Middle East. Elsewhere in Africa, Asia, Australia and North America, it has been introduced as a food fish or as a control of aquatic vegetation. Where introduced, it sometimes becomes invasive, threatening the local ecology and species. The redbelly tilapia is an important food fish and sometimes aquacultured.
Mayaheros is a genus of cichlid fish that is native to Mexico and northern Central America. This genus has a disjunct distribution, with the M. urophthalmus group being found in the Atlantic drainages of southeastern Mexico, Belize, eastern Guatemala, northern Honduras and northeastern Nicaragua, while M. beani is the northernmost cichlid in the Pacific drainage, ranging from Jalisco to Sonora in northwestern Mexico. Both inhabit a wide range of habitats such as freshwater rivers, streams, lakes, ponds, pools and marshes, as well as brackish waters like estuaries, lagoons and mangrove, with members of the M. urophthalmus group sometimes even occurring in caves or coastal marine waters. They are medium-sized to large cichlids that are omnivorous, feeding mostly on small animals, but also plants and detritus.
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