The Carboniferous is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period 358.9 million years ago (mya), to the beginning of the Permian Period, 298.9 mya. The name Carboniferous means "coal-bearing", from the Latin carbō ("coal") and ferō, and refers to the many coal beds formed globally during that time.
The Permian is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia.
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthparts adapted to rasping or grinding. Horses and other herbivores have wide flat teeth that are adapted to grinding grass, tree bark, and other tough plant material.
Diadectidae is an extinct family of early tetrapods that lived in what is now North America and Europe during the Late Carboniferous and Early Permian, and in Asia during the Late Permian. They were the first herbivorous tetrapods, and also the first fully terrestrial animals to attain large sizes. Footprints indicate that diadectids walked with an erect posture. They were the first to exploit plant material in terrestrial food chains, making their appearance an important stage in both vertebrate evolution and the development of terrestrial ecosystems.
Edaphosaurus is a genus of extinct edaphosaurid synapsids that lived in what is now North America and Europe around 303.4 to 272.5 million years ago, during the Late Carboniferous to Early Permian. American paleontologist Edward Drinker Cope first described Edaphosaurus in 1882, naming it for the "dental pavement" on both the upper and lower jaws, from the Greek edaphos έδαφος and σαῦρος ("lizard").
Edaphosauridae is a family of mostly large Late Carboniferous to Early Permian synapsids. Edaphosaur fossils are so far known only from North America and Europe.
Eupelycosauria is a large clade of animals characterized by the unique shape of their skull, encompassing all mammals and their closest extinct relatives. They first appeared 308 million years ago during the Early Pennsylvanian epoch, with the fossils of Echinerpeton and perhaps an even earlier genus, Protoclepsydrops, representing just one of the many stages in the evolution of mammals, in contrast to their earlier amniote ancestors.
Xenacanthida is a super-order of extinct shark-like chondrichthyans known from the Carboniferous to Triassic. They were native to freshwater, marginal marine and shallow marine habitats. Some xenacanths may have grown to lengths of 5 m (16 ft). Most xenacanths died out at the end of the Permian in the End-Permian Mass Extinction, with only a few forms surviving into the Triassic.
Bandicota is a genus of rodents from Asia known as the bandicoot rats. Their common name and genus name are derived from the Telugu language word pandikokku (పందికొక్కు). DNA studies have found the group to be a monophyletic clade sister to the radiation of Molucca and Australian Rattus species as part of the paraphyletic Rattussensu lato.
Caseidae are an extinct family of basal synapsids that lived from the Late Carboniferous to Middle Permian between about 300 and 265 million years ago. Fossils of these animals come from the south-central part of the United States, from various parts of Europe, and possibly from South Africa if the genus Eunotosaurus is indeed a caseid as some authors proposed in 2021. Caseids show great taxonomic and morphological diversity. The most basal taxa were small insectivorous and omnivorous forms that lived mainly in the Upper Carboniferous and Lower Permian, such as Eocasea, Callibrachion, and Martensius. This type of caseid persists until the middle Permian with Phreatophasma and may be Eunotosaurus. During the early Permian, the clade is mainly represented by many species that adopted a herbivorous diet. Some have evolved into gigantic forms that can reach 6–7 metres (20–23 ft) in length, such as Cotylorhynchus hancocki and Alierasaurus ronchii, making them the largest Permian synapsids. Caseids are considered important components of early terrestrial ecosystems in vertebrate history because the numerous herbivorous species in this family are among the first terrestrial tetrapods to occupy the role of primary consumer. The caseids experienced a significant evolutionary radiation at the end of the early Permian, becoming, with the captorhinid eureptiles, the dominant herbivores of terrestrial ecosystems in place of the edaphosaurids and diadectids.
Ianthasaurus is an extinct genus of small edaphosaurids from the Late Carboniferous.
Otsheria is an extinct genus of anomodont, in the infraorder venyukovioidea. It lived in modern-day Russia during the Permian.
Barameda is a genus of rhizodont lobe-finned fishes which lived during the Tournaisian stage near the start of the Carboniferous period in Australia; fossils of the genus have been reported from the Snowy Plains Formation. The largest member of this genus, Barameda decipiens, reached an estimated length of around 3–4 metres (9.8–13.1 ft), while smallest species, B. mitchelli is estimated to have had a length of about 35 centimetres (14 in).
Letognathus is a genus of rhizodont tetrapodomorph that lived during the Carboniferous period. Its remains come from the Blue Beach Member of the Horton Bluff Formation, near Hantsport, Nova Scotia. Like most rhizodonts, it was of relatively large size, had a large recurved fang at the symphysis of the lower jaw, and a row of three coronoid fangs along the length of the jaw in addition to its marginal dentition. Letognathus is important for rhizodont systematics because it retains a number of primitive features, such as ossified Meckel's cartilage, are not found in the genera Rhizodus and Strepsodus.
Glaucosaurus is an extinct genus of small edaphosaurids from the Early Permian. The type species, G. megalops, was named in 1915.
Macromerion is an extinct genus of non-mammalian synapsids, specifically Pelycosaurs, in the family Sphenacodontidae from Late Carboniferous deposits in the Czech Republic. It was named as a species of Labyrinthodon in 1875 and as its own genus in 1879.
Copelatus sociennus is a species of diving beetle. It is part of the subfamily Copelatinae in the family Dytiscidae. It was described by J. Balfour-Browne in 1952. The species can be found on Coloane and in Seac Pai Van. It feeds on Nelumbo nucifera.
Occidens is an extinct genus of stem tetrapod that lived during the earliest part of the Carboniferous in what is now Northern Ireland. It is known from a single type species, Occidens portlocki, named in 2004 on the basis of a left lower jaw described by British geologist Joseph Ellison Portlock in 1843.
NGC 4459 is a lenticular galaxy located about 50 million light-years away in the constellation of Coma Berenices. NGC 4459 is also classified as a LINER galaxy. NGC 4459 was discovered by astronomer William Herschel on January 14, 1787. NGC 4459 is a member of the Virgo Cluster.
Gordodon is an extinct genus of non-mammalian synapsid that lived during the Early Permian of what is now Otero County, New Mexico. It was a member of the herbivorous sail-backed family Edaphosauridae and contains only a single species, the type species G. kraineri. Gordodon is unusual among early synapsids for its teeth, which were arranged similarly to those of modern mammals and unlike the simple, uniform lizard-like teeth of other early herbivorous synapsids. Gordodon had large incisor-like teeth at the front, followed by a prominent gap between them and a short row of peg-like teeth at the back. Gordodon was also relatively long-necked for an early synapsid, with elongated and gracile vertebrae in its neck and back. Like other edaphosaurids, Gordodon had a tall sail on its back made from the bony neural spines of its vertebrae. The spines also had bony knobs on them, a common trait of edaphosaurids, but the knobs of Gordodon are also unique for being more slender, thorn-like and randomly arranged along the spines. It is estimated to have been rather small at 1 m in length excluding the tail and 34 kg (75 lb) in weight.
